A new marattiaceous fern from the Lower Jurassic of Patagonia (Argentina): the renaissance of Marattiopsis

A new marattiaceous fern from the Lower Jurassic of Patagonia (Argentina): the renaissance of Marattiopsis
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DOI:
10.1080/14772019.2014.936974
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发表时间:
2015-08-03
影响因子:
2.6
通讯作者:
Scasso, Roberto
Scasso, Roberto
中科院分区:
地球科学2区
文献类型:
--
作者:
Escapa, Ignacio H.;Bomfleur, Benjamin;Scasso, Roberto

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我们在这里描述Marattiopsis patagonica sp. nov.(Marattiaceae)的营养和肥沃的叶子的印象,从下侏罗纪的丘布特省,巴塔哥尼亚,阿根廷。该新种具有独特的营养和生殖特征,包括一个突出的基叶耳;细波状,具小齿,到有细锯齿的羽片边缘;间质纤维(小静脉返流)与规则静脉互生;合囊的近边缘位置和独特的相对长度(约为小羽片宽度的12-20%);每个合囊的孢子囊对数量相对较少(通常为14或16)。新物种被分配到形态属Marattiopsis,因为它没有显示出任何特征,可以明确地放置在任何现代属,Marattias.s.,Ptisana或Eupodium。新成立的paraphyly的Marattias.l.物种对化石物种的分类有着深远的影响。在这种情况下,我们提供了一个简要的评估的意义的化石Marattiopsis系统发育和地理学的解释。看来,基本上是现代的marattioid蕨类植物,包括Marattiopsis,这通常被认为是典型的元素,在北方热带地区的中生代,可能留下了重要的,但在很大程度上被忽视的化石记录在亚热带冈瓦纳。
We here describe Marattiopsis patagonica sp. nov. (Marattiaceae) based on vegetative and fertile foliage impressions from the Lower Jurassic of Chubut Province, Patagonia, Argentina. The new species exhibits a unique combination of vegetative and reproductive features, including a prominent basal auricle; finely undulate, denticulate, to serrulate pinnule margins; interstitial fibres (venuli recurrentes) alternating with regular veins; submarginal position and distinctive relative length of synangia (about 12-20% of the pinnule width); and relatively small number of sporangia pairs per synangium (usually 14 or 16). The new species is assigned to the morphogenus Marattiopsis, since it shows no characters that would allow an unambiguous placement in any of the modern genera, Marattias.s., Ptisana, or Eupodium. The newly established paraphyly of the Marattias.l. species has a profound impact on the classification of fossil species. In this context, we provide a brief assessment of the significance of the fossil Marattiopsis for phylogenetic and biogeographical interpretations. It appears that essentially modern marattioid ferns, including Marattiopsis, which are commonly considered to be typical elements of tropical areas of the northern hemisphere during the Mesozoic, may have left an important but largely ignored fossil record in subtropical Gondwana.