The opening and shedding mechanism of the female cones of Pinus radiata
The opening and shedding mechanism of the female cones of Pinus radiata
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DOI：
10.1071/bt9640125
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发表时间：
1964-01-01
期刊：
影响因子：
--
通讯作者：
WARDROP, A. B.
中科院分区：
文献类型：
--
作者：
ALLEN, R.;WARDROP, A. B.
研究表明，放射草雌球果的开放是由于球果鳞片的近轴维管组织和远轴厚壁组织的差异收缩所致。管胞次生壁的组织通常由三个螺旋组织的同心层组成。在外层和内层，微纤维取向大致为横向，中层螺旋与细胞纵轴成约40度角。在厚壁组织中，次生壁由宽层组成，其中薄片的微纤丝几乎横跨细胞轴，与窄层交替，薄片的微纤丝几乎平行于细胞轴。在开放之前，通过用树脂封闭花梗管胞的管腔，切断圆锥体与茎或枝之间的血管连接。随着茎的径向生长，茎或枝的木质部和球果花序梗的木质部之间会产生小裂缝。裂隙充满树脂，花序梗的管胞逐渐受到侵蚀，直到花序梗的木质部与茎或枝的木质部最终分离。|抽象作者：作者
The opening of the female cones of P. radiata has been shown to result from differential shrinkage between the adaxial vascular tissue and the abaxial sclerenchyma of the cone scale. The organization of the secondary wall of the tracheids typically consists of three helically organized concentric layers. In the outer and inner layers the micro-fibrillar orientation is approximately transverse, and in themiddle layer the helix makes an angle of c. 40[degree] with the longitudinal cell axis. In the sclerenchyma the secondary wall consists of wide layers in which the microfibrils of the lamellae are almost transverse to the longitudinal cell axis, alternating with narrow layers in which the microfibrils of the lamellae are almost parallel to the cell axis. Opening is preceded by a severance of the vascular connection between the cone and the stem or branch by the occlusion of the lumina of the tracheids of the peduncle with resin. As radial growth of the stem, proceeds, small fissures develop between the xylem of the stem or branch and that of the cone peduncle. The fissures become filled with resin and there is a progressive erosion of the tracheids of the peduncle until ultimately the xylem of the peduncle is separated from that of the stem or branch. || ABSTRACT AUTHORS: Authors