The genetics of corky; the New World alleles and their possible role as an interspecific isolating mechanism.

The genetics of corky; the New World alleles and their possible role as an interspecific isolating mechanism.
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DOI:
10.1007/bf02986784
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发表时间:
1946
影响因子:
1.5
通讯作者:
Stephens Sg
Stephens Sg
中科院分区:
生物学4区
文献类型:
--
作者:
Stephens Sg

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[~NTRODUOTION Harland(]915,I939)在跨越某些波旁威士忌(~oxsjph~‘m./lt:<sv.)时首次被注意到。H~M~VAT。R~,c~‘ie-gcda~,te}futohinson)。H海岛卵石(bcorDac2e~‘e L.)所获得的Hyb:Dd是一株正常的、未取食的植物,叶呈黄色斑点,茎S,~-M~+粗CO。Rkv OUP,‘owt,h’将异常的‘corky’bqJe与g.ba.~bc1,de)ZSE进行重复杂交,得到明显的1:Corky‘:1正常分离,这表明Conplen:ent:Ary基因是导致木栓异常产生的原因。他暂时将它们分别命名为Ck P Ck(波旁带)和Ck P Gk S(海岛带)。从t9:39开始,收集到的数据并不能证实哈兰德的解释。埃里奇,但建议使用dmt。ABNOJ:MAL COR]x是丹对互作o:[等位基因在单个座位上,即所有科基计划~S都是杂合子]。由于轮胎复杂的女性不孕因素、改性剂复合体的不同效应以及对环境条件的不同反应,临界分析被证明是令人惊讶的不同因素。本文旨在说明在过去的一年中软木生产的异常。E~SSE x~zm‘~:E-Gek,.’~‘~Ze杂交种,并考虑其发生的地理分布。由于软木塞的产生只是复杂异常的一个征兆,因此在开头用R来描述表现为遗传异常的异常性状将是方便的。与正常外观的第一个偏差是在开花之前],有时早到冷杉叶。叶片在叶脉附近变黄,斑块中的叶绿素变得‘干净’。淡黄色的斑驳的淡啤酒铺满了整个区域。整片叶子。叶片的边缘倾向于向内滚动,因此GHA~~叶表面变成槽形,而不是FLA~。在中,催吐论被缩短了,而I~极端情况下~他、计划~的主轴被贪婪地压缩,使得I~变得矮小。WRH拥挤的树叶。这是最矮小的一种。开花前开花。此外,还有一个缺点:由于产量过剩,侧枝增加了营养部分的拥挤外观。软木塞的产量变化很大,当植物被遮蔽或生长在异常潮湿的环境中时,软木产量往往会受到抑制。在正常田间条件下,高级木栓型的整个茎、贝里克里和叶中脉都覆盖着厚厚的灰色软木层。软木塞通常第一次出现时是沿着仍在迅速伸长的茎节间的细纹r(Hming Longitudina);因此,与正常的软木塞生产时间相比,R出现得更早。在低品位类型中,可能只有少量的软木;
].~ NTRODUOTION Harland (] 915, I939) first noted in crossing certain 8ypes of Bourbon (~ oxsjph~'m./Lt:< sv. h~ m~ vat. r~, c~'ie-gcda~, te} futohinson) wit. h Sea Island ooSton (g. bcorDac2e~'e L.) tha~ the hyb: dd obtained was an~ d) nonnal st, unfed plant wRh leaves< showing yellow spotting of the: _,.~ os~ ic type and the stem s,,~-M~ ed with~ rough co. rkv ouP,'owt, h Repeated b~ e!<-crossing of the abnormal'corky'bqJe to g. ba.~ bcl, de) zse gave clear 1: corky': 1 normal segregations, which suggested that con: plen: ent: ary genes were responsible for abnormal cork production. These he named provisionally Ck P ck (carried by Bourbon) and ck P Gk s (carried by Sea island) respectively. Since t9: 39 data have been collected which do not confirm Harland's interpret. erich but suggest dmt. abnoJ: mal cor] x is dan to the interaction o:[alleles at a single locus, ie that all corky plan~ s are heterozygotes. Critical analysis has proved to be surprisingly diflqcuR owing to tire complicating factors of female sterility, differential effects of modifier complexes and di: fferen~ ial response to environmental conditions, it is the p'arpose of this paper to give an account of abnormal cork production in oer~, ain ba'rbe~-~.? e~ sse x~ zm'~: e-gek,.'~'~ Ze hybrids, and to consider the geographic clistribution of its occurrence.2. T~ s com< Y CO>~ PLEX Since cork production is only one symptom of a complex abnormality, R will be convenien~ at the outset f, o describe more fntly the abnormal characters which behave as a genetic unR. The first deviation from the normal appearance is seen] prior to flowering, sometimes as early as the firs% foliage leaf. The leaves become yellow in the neighbot~ rhood of the veins, and patches become'cleared'of chlorophyll. Lager a yellowish mottling spreads over the. whole leaf. The edges of the leaves tend to roll inwards so gha~~ he leaf surface becomes trough-shaped instead of fla~. In, emetics are shortened, and i~ extreme cases the main axis of~ he, plan~ is greedy cotnpressed so that i~ becomes dwarfed. wRh crowded leaves. This dwarfing is most nogicea. ble prior~ o flowering. Also there is a~ endency: for an excess production, of lateral branches which enhances the crowded appearance of the vegetative parts. Cork production is very variable and is often suppressed when the plants are shaded or grown under exceptiotmlty humid condbions. Under normal field conditions high-grade corky types h~ ve the whole of the stems, pericles and leaf midribs covered with~ thick greyish layer of cork. The cork usually makes its first appearance as thin streaks r (hming longitudina] ly along the still rapidly elongating stem internodes; hence R occurs prema~ u. rely in comparison xvigk the time of normal cork production. In low grade types only fain {traces of cork may be apparen<