Pachybolini -: a tribe of giant Afrotropical millipedes:: arguments for monophyly and the description of a new genus from Madagascar (Diplopoda: Spirobolida: Pachybolidae)

Pachybolini -: a tribe of giant Afrotropical millipedes:: arguments for monophyly and the description of a new genus from Madagascar (Diplopoda: Spirobolida: Pachybolidae)
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DOI:
10.1071/is07008
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发表时间:
2008-01-01
影响因子:
2.2
通讯作者:
Waegele, Johann-Wolfgang
Waegele, Johann-Wolfgang
中科院分区:
生物学2区
文献类型:
--
作者:
Wesener, Thomas;Enghoff, Henrik;Waegele, Johann-Wolfgang

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Madabolus,gen. nov. maximus,sp. nov.(模式种),分布于马达加斯加西部的几个地区。根据形态特征,对26个螺菌分类群进行了系统发育分析,将Madabolus,gen. nov.与非洲热带属Pelmatojulus de Saussure,1860; Pachybolus Cook,1897; Hadrobolus Cook,1897和Epibolus Cook,1897。Pachybolini族包括Madabolus,gen. nov.和上述四个属。Pachybolini的假定自形是:一个封闭的切迹外侧;颚chilarial板舌顶端的刚毛;外阴肾形或新月形,与阀门会议在一个弯曲的裂缝和嵴突出。在Pachybolini内,Pelmatojulus(西非)凭借顶端触角感觉锥数量的增加而脱颖而出,很有可能是一种无定形。Madabolus,gen. nov.和东非属的部落共享另一个潜在的synapomorphy,即,刚毛数量增加的gnathochilarial lamelae linguales。其他马达加斯加属的Pachybolidae似乎不相关的非洲热带属。
Madabolus, gen. nov. is described to accommodate M. maximus, sp. nov. (type species) from several localities in western Madagascar. A phylogenetic analysis of 26 spirobolidan taxa, based on morphological characters, places Madabolus, gen. nov. with the Afrotropical genera Pelmatojulus de Saussure, 1860; Pachybolus Cook, 1897; Hadrobolus Cook, 1897 and Epibolus Cook, 1897. The tribe Pachybolini is proposed to include Madabolus, gen. nov. and the four abovementioned genera. Putative autapomorphies of Pachybolini are: a closed incisura lateralis; setae of gnathochilarial lamella lingualis apical; vulva kidney- or crescent-shaped, with valves meeting in a sinuous fissure and crest protruding. Within Pachybolini, Pelmatojulus (West Africa) stands out by virtue of an increased number of apical antennal sensory cones, with great probability an apomorphy. Madabolus, gen. nov. and east African genera of the tribe share another potential synapomorphy, namely, an increased number of setae on the gnathochilarial lamellae linguales. Other Malagasy genera of Pachybolidae do not seem to be related to Afrotropical genera.