A new species of Pachynolophus (Mammalia, Perissodactyla) from Zambrana (Alava, Basque-Cantabrian Region).: Phylogenetic analysis of Pachynolophus and first mention in the Late Eocene of the Iberian Peninsula
A new species of Pachynolophus (Mammalia, Perissodactyla) from Zambrana (Alava, Basque-Cantabrian Region).: Phylogenetic analysis of Pachynolophus and first mention in the Late Eocene of the Iberian Peninsula
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DOI：
10.1016/j.geobios.2003.07.005
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发表时间：
2005-01-01
期刊：
影响因子：
1.6
通讯作者：
Cuesta, MA
中科院分区：
文献类型：
--
作者：
Badiola, A;Pereda-Suberbiola, X;Cuesta, MA
赞布拉纳始新世晚期的最新野外工作（Alava，Basque-Cantabrian Region）发现了一个新的厚头类物种：P. zambranensis nov. sp.根据本研究中的系统发育分析，Zambrana种是单系厚头类的一部分，至少由三种联会形态定义：M1/1-M3/3系列的尺寸明显递增，M3/3明显大于M2/2; M3/相对于M2/表面的相对表面适度增加，下部系列唇扣带回较弱。Pachynolophus包括种P duvali、P. livinierensis、P. garimondi、P. lavocati。桑布拉尼亚变形杆菌、波依达变形杆菌和切塞拉斯变形杆菌。P. molipotensis和P. cayluxi没有足够的材料将它们归属于Pachynolophus。P. bretovensis不具有Pachynolophus的突触形态，P. hookeri不应被称为该属。P. zambranensis与P. garimondi和P. lavocati有几个共同的形态特征，即：P3-4/系列的后原嵴是一个明确的lophe; P4/具有近四边形的轮廓，并显示出强大的后舌带，形成后部增宽;有一个“初期的中柱”（pseudoestyle？）在上臼齿上P. zambranensis的主要特征是脊齿，在P3-4/系列中，后原嵴斜向外嵴，在P3-4/系列中也有一个“假柱”。它代表了在伊比利亚半岛始新世晚期描述的Pachynolophus属的首次提及。(C)2005年爱思唯尔SAS。All rights reserved.
Recent fieldwork in the Late Eocene of Zambrana (Alava, Basque-Cantabrian Region) has yielded a new species of Pachynolophils: P. zambranensis nov. sp. According to the phylogenetic analysis made in this study, the Zambrana species is part of the monophyletic Pachynolophus, which is defined at least by three synapomorphies: significant progressive increase of the size in the M1/1-M3/3 series, being M3/3 outstandingly bigger than M2/2; moderate increase of the relative surface of M3/ with respect to M2/ surface, and weak labial cingula in the lower series. Pachynolophus includes the species P duvali, P. livinierensis, P garimondi, P. lavocati. P zambranensis, P boixedatensis and P. cesserasicus. P. molipotensis and P. cayluxi are not represented by sufficient material to attribute them to Pachynolophus. P. bretovensis does not share a synapomorphy of Pachynolophus and P. hookeri should not be referred to this genus. P. zambranensis shares several synapomorphies with P. garimondi and P. lavocati, which are: the posprotocrista,of the P3-4/ series is a well-defined lophe; the P4/ has a subquadrangular outline and shows a strong posterolingual cingulum, forming a posterior widening; and there is an "incipient mesostyle" (pseudomesostyle?) in the upper molars. The species P. zambranensis is mainly characterized by lophodont dentition, with the postprotocrista obliquely oriented to the ectoloph in the P3-4/ series, and a "pseudomesostyle" also present in the P3-4/ series. It represents the first mention of the genus Pachynolophus described in the Late Eocene of the Iberian Peninsula. (C) 2005 Elsevier SAS. All rights reserved.