THE BOMBYCOIDEA - PHYLOGENY AND HIGHER CLASSIFICATION (LEPIDOPTERA, GLOSSATA)
THE BOMBYCOIDEA - PHYLOGENY AND HIGHER CLASSIFICATION (LEPIDOPTERA, GLOSSATA)
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DOI：
10.1163/187631294x00045
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发表时间：
1994-01-01
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影响因子：
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通讯作者：
MINET, J
中科院分区：
文献类型：
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作者：
MINET, J
本文主要旨在重新评估四个蚕科科的界限，即蚕蛾科、蚕蛾科、柠檬蛾科和婆罗门蛾科。为整个“bombycoid 复合体”（Mimallonoidea + Lasiocampoidea + Bombycoidea）提出了一个不完全解析的分支图。在 Bombycoidea 中，主要的二分法被认为位于 Eupterotidae + Bombycidae s.lat 之间。 + Endromidae + Mirinidae + Saturniidae，以及 Carthaeidae + Lemoniidae + Brahmaeidae + Sphingidae。 Lemoniidae 和 Brahmaeidae 具有至少九个同源性，被认为是可靠的姐妹类群，并且 Lemoniidae + Brahmaeidae 被提议作为天蛾科的姐妹类群。另一个新提出的进化枝将 Endromidae、Mirinidae 和 Saturniidae 归为一类。在科一级，Hibrildidae 与 Eupterotidae (syn. n.) 是同义的，其最重要的自体变态在于先前未被注意到的雌性后腿的特殊性（二跗节通常具有腹中排的刺）。腿部结构的性别二态性也导致了对Saturniidae的重新定义，该科必须包括现代作家的“Oxytenidae”和“Cercophanidae”，其亚科等级（stat.rev.）。因此，雌性前腿第四跗节上的一对远端齿状结构以及其他一些令人信服的自体特征可以归因于土星科的地面平面。另一方面，“Apatelodidae”只是暂时被归入“Bombycidae sensu lato”，这是一个在 Apatelodidae sensu auct 范围内暂时复活的群体。证明是双系的。事实上，考虑到相当多的共源性，“apatelodid”亚科 Epiinae 与 Bombycinae (syn. n.) 同义。当进行更广泛的研究时，雄性腹部第八胸骨的形态可能会导致蚕科的概念略有不同，更受限制（即排除“真正的”Apatelodidae）。通常被认为是不确定的物种，非洲 Sabalia Walker 属和 Spiramiopsis Hampson 明确分别属于 Lemoniidae 和 Brahmaeidae。记录并讨论了这两个科的自体变态，以及天蛾科的特征。后者暂时被认为有 3 个亚科（Smerinthinae stat. rev.、Sphinginae、Macroglossinae），Eupterotidae 中有 5 个亚科（Hibrildinae、Tissanginae、Janinae、Panacelinae、Eupterotinae），以及 Bombycidae s.lat 中有 4 个亚科。 （Apatelodinae、Phiditiinae subfam. n.、Prismostictinae [= Oberthueriinae, syn. rev.]、Bombycinae）。其中三个亚科被认为是新意义上的，即。 Panacelinae、Apatelodinae 和 Bombycinae。尽管本研究没有充分考虑土星科的系统发育，但对土星科的三个亚科（Oxyteninae、Cercophaninae、Ludiinae）的组成进行了严格的研究，并且土星亚科被视为该科中最“原始”的成员。
This paper is chiefly aimed at reassessing the limits of four bombycoid families, namely the Eupterotidae, Saturniidae, Lemoniidae, and Brahmaeidae. An incompletely resolved cladogram is proposed for the whole 'bombycoid complex' (Mimallonoidea + Lasiocampoidea + Bombycoidea). Within the Bombycoidea, the primary dichotomy is considered to lie between the Eupterotidae + Bombycidae s.lat. + Endromidae + Mirinidae + Saturniidae, and the Carthaeidae + Lemoniidae + Brahmaeidae + Sphingidae. Sharing at least nine synapomorphies, the Lemoniidae and Brahmaeidae are regarded as reliable sister groups, and the Lemoniidae + Brahmaeidae are proposed as a sister group to the Sphingidae. Another newly proposed clade groups together the Endromidae, Mirinidae and Saturniidae. At family level, the Hibrildidae are synonymized with the Eupterotidae (syn. n.), for which the most significant autapomorphy lies in a previously unnoticed particularity of the female hind leg (distitarsus typically provided with a midventral row of spines). Sexual dimorphism in leg structure also leads to a redefinition of the Saturniidae, a family which must include, with subfamily rank (stat. rev.), the 'Oxytenidae' and 'Cercophanidae' of modern authors. A pair of distal, tooth-like structures on the fourth tarsomere of the female fore leg can thus be ascribed to the ground plan of the Saturniidae, along with a few other convincing autapomorphies. On the other hand, the 'Apatelodidae' are only tentatively placed in the 'Bombycidae sensu lato', a group provisionally resurrected insofar as the Apatelodidae sensu auct. prove to be diphyletic. As a matter of fact, the 'apatelodid' subfamily Epiinae is synonymized with the Bombycinae (syn. n.) in consideration of a rather large number of synapomorphies. When more extensively studied, the morphology of the eighth sternum of the male abdomen might lead to a slightly different, more restricted, concept of the Bombycidae (i.e. excluding 'true' Apatelodidae). Often regarded as incertae sedis, the African genera Sabalia Walker and Spiramiopsis Hampson are definitely assigned to the Lemoniidae and Brahmaeidae respectively. Autapomorphies of these two families are recorded and discussed, as are those found to characterize the Sphingidae. Three subfamilies are tentatively recognized within the latter (Smerinthinae stat. rev., Sphinginae, Macroglossinae), five within the Eupterotidae (Hibrildinae, Tissanginae, Janinae, Panacelinae, Eupterotinae), and four within the Bombycidae s.lat. (Apatelodinae, Phiditiinae subfam. n., Prismostictinae [= Oberthueriinae, syn. rev.], Bombycinae). Three of these subfamilies are considered in a new sense, viz. the Panacelinae, Apatelodinae, and Bombycinae. Although the phylogeny of the Saturniidae is not fully taken into account in the present study, the composition of three saturniid subfamilies is critically examined (Oxyteninae, Cercophaninae, Ludiinae), and the Oxyteninae are viewed as the most 'primitive' member of the family.