Molecular phylogenetic analyses and systematics of the genus Saussurea and related genera (Asteraceae, Cardueae)
Molecular phylogenetic analyses and systematics of the genus Saussurea and related genera (Asteraceae, Cardueae)
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作者：
Kita, Y;Fujikawa, K;Kato, M
标题风毛菊属的系统发育关系。利用核糖体ITS和叶绿体matK基因分析了印度棉（Hemistepta lyrata Bunge）和印度棉（Hemistepta lyrata Bunge）的基因型。系统发育分析表明，角果风毛菊（Saussurea cerato-carpa Decne.）的subg。Jurinocera（Baill.）利普施并将该种归入Elatae Hook. f.组。的subg。风毛菊属（Saussurea）是从风毛菊属（Saussurea）的所有其它成员中分离出来的。亚属的种类。Jurinocera和亚属（subgen.）风毛菊属Elatae组沿着包括Jurismus卡斯在内的一个组形成一个单系单位。Subg.的种类。Jurinocera和Sect. Elatae共享4-5-cypselas和顶端果皮冠，带有尖角或锋利的牙齿。风毛菊该小组由该小组六个科的代表组成。风毛菊属（Aussurea）和形态特征独特的亚属[Amphilaena（Stschegl.）利普施，Eriocoryne（DC.）胡克·F，和西奥多莉亚（卡斯。）Lipsch.]。两种适应寒冷的形式，亚属的“雪球植物”。Eriocoryne和亚属的“温室植物”。Amphilaena，被确认为风毛菊属的成员。琴叶半步甲和风毛菊显示出一个密切的姐妹关系，这是由冠毛二型性支持的，涉及离生，短，粗糙的外部刚毛或微小的外部鳞片，基部合生在一个环中，具有更长，羽状的内部刚毛。
The phylogenetic relationships of the genus Saussurea DC. and Hemistepta lyrata Bunge were analyzed using the nuclear ribosomal ITS and chloroplast matK gene. Phylogenetic analyses revealed that Saussurea cerato-carpa Decne. of subg. Jurinocera (Baill.) Lipsch. and the species assigned to sect. Elatae Hook.f. of subg. Saussurea were separated from all other members of the genus Saussurea. The species of subgen. Jurinocera and subgen. Saussurea sect. Elatae form a monophyletic unit along with a group including genus Jurinea Cass. The species of subg. Jurinocera and sect. Elatae share 4-5-gonal cypselas and apical pericarpal crowns bearing sharp horns or sharp teeth. The Saussurea s.s. group is comprised of representatives of six sections of the subg. Saussurea and morphologically distinctive subgenera [Amphilaena (Stschegl.) Lipsch., Eriocoryne (DC.) Hook.f., and Theodorea (Cass.) Lipsch.]. Two frigid-adapted forms, the "snowball plants" of subg. Eriocoryne and the "glasshouse plants" of subg. Amphilaena, were confirmed as members of Saussurea s.s. Hemistepta lyrata and Saussurea s.s. show a close sister relationship, which is supported by the pappus dimorphism involving free, short, scabrid outer bristles or tiny outer scales and basally connate in a ring, with much longer, plumose inner bristles.