The Late Carboniferous Triplosoba pulchella is not a fly in the ointment but a stem-mayfly
The Late Carboniferous Triplosoba pulchella is not a fly in the ointment but a stem-mayfly
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DOI：
10.1111/syen.12103
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发表时间：
2015-04-01
影响因子：
4.8
通讯作者：
Bethoux, Olivier
中科院分区：
文献类型：
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作者：
Bethoux, Olivier
晚石炭世三虫（Triplosoba pulchella, Brongniart, 1893）被大多数作者认为是一种茎蝇。与此相反，在该物种的最新修订中，它与已灭绝的古翼目大翼目有关。该物种唯一可用的标本使用各种人工和计算机辅助光学技术重新调查。提出新的数据并与以前的数据进行比较。由于观察不完全，“翼龙假说”的推理是有缺陷的。此外，这一假设还需要在蛇翼亚纲中存在多重角色冲突。相反，“茎-蜉蝣假说”更受青睐。在pulchella的一个后翅中观察到一个分叉的RP2 （RP2在另一个后翅中是简单的，在前翼中保存不完全）是该群体的严格的形像。它还表明，该物种拥有什么被解释为一个早期状态的肋撑。其他的观察结果并不与这种定位相矛盾，总的来说，这种定位比把它归为蝶亚目更加一致。这一贡献表明了长期保护自然历史收藏品的重要性，使新的调查技术在未来得到应用。
The Late Carboniferous Triplosoba pulchella (Brongniart, 1893) was regarded by most authors as a stem-mayfly. Contrary to this, it was related to the extinct palaeopteran' order Diaphanopterodea in the most recent revision of the species. The only available specimen of the species is re-investigated using various manual and computer-assisted optical techniques. New data are presented and compared with previous data. The reasoning underlying the Diaphanopterodea hypothesis' is found to be deficient due to incomplete observations. Moreover, this hypothesis requires multiple character conflicts within Diaphanopterodea. Instead, the (stem-)Ephemeroptera hypothesis' is favoured. The occurrence of a forked RP2, observed in one hindwing of T. pulchella (RP2 is simple in the other hindwing, and incompletely preserved in forewings) is a strict apomorphy of the group. It is also shown that the species possessed what is interpreted as an early state of the costal brace. Other observations do not contradict this placement, which is overall more consistent than an assignment to the Diaphanopterodea. This contribution demonstrates the importance of long-term conservation of natural history collections, allowing new investigation techniques to be applied in the future.