PALAEOBIOGEOGRAPHY AND EVOLUTIONARY PATTERNS OF THE LARGER FORAMINIFER BORELIS DE MONTFORT (BORELIDAE)
PALAEOBIOGEOGRAPHY AND EVOLUTIONARY PATTERNS OF THE LARGER FORAMINIFER BORELIS DE MONTFORT (BORELIDAE)
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DOI：
10.1002/spp2.1273
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发表时间：
2021-02-01
影响因子：
2.3
通讯作者：
Iryu, Yasufumi
中科院分区：
文献类型：
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作者：
Bassi, Davide;Braga, Juan C.;Iryu, Yasufumi
alveolinoid Borelis物种的古生物地理学揭示了导致两个现存代表的进化模式，这两个代表发生在浅水热带碳酸盐岩，珊瑚礁相关的设置。通过对Borelis化石种的模式标本和新标本的研究，沿着对Borelis化石种的分类地位、种围（根据前房大小、Y形隔的出现和伸长指数）、古生物地理和演化动力学进行了评价。这里讨论的物种仅从化石（B. pygmaea，B. B. inflata，B. philippinensis，B.梅洛，B。curdica），并从化石和现代（B. pulchra，B. Schlumbergeri）标本。第一次，化石和最近Borelis标本说明通过微型计算机断层扫描图像。根据Y形隔膜的出现，两个谱系被区分开来。第一谱系包括B类，起源于中-上始新世Borelis vonderschmitti。inflata，B. pulchra和B.矮缩，缺乏Y形小隔。第一个具Y形隔片的种是鲁佩利亚种B。产于印度-太平洋西部的菲律宾珊瑚。B的西迁者。philippinensis进入地中海后产生了B。梅洛（Aquitanian-Messinian）和B. curdica（Burdigalian-Tortonian）.这两个物种由于朗基期地中海盆地的东部封闭而与印度太平洋隔离，并在墨西拿盐度危机期间消失。从托顿人开始，B。schlumbergeri，起源于B. philippinensis，与B一起沿着居住在印度-太平洋地区。pulchra。来自太平洋中部的B。早在Piacenzian关闭中美洲航道之前，pulchra就到达了加勒比地区。
The palaeobiogeography of the alveolinoid Borelis species reveals the evolutionary patterns leading to the two extant representatives, which occur in shallow-water tropical carbonate, coral reef-related settings. Type material and new material of fossil Borelis species, along with Recent specimens were studied to assess their taxonomic status, species circumscriptions (based on proloculus size, occurrence of Y-shaped septula, and the index of elongation), palaeobiogeography and evolutionary dynamics. The species dealt with here are known from exclusively fossil (B. pygmaea, B. inflata, B. philippinensis, B. melo, B. curdica), and from fossil and modern (B. pulchra, B. schlumbergeri) specimens. For the first time, fossil and Recent Borelis specimens are illustrated via micro-computed tomography scanning images. Depending on the occurrence of Y-shaped septula, two lineages are distinguished. Deriving from the middle-upper Eocene Borelis vonderschmitti, the first lineage includes B. inflata, B. pulchra and B. pygmaea, lacking Y-shaped septula. The first species bearing Y-shaped septula is the Rupelian B. philippinensis of the western Indo-Pacific. The westward migrants of B. philippinensis into the Mediterranean gave rise to B. melo (Aquitanian-Messinian) and B. curdica (Burdigalian-Tortonian). These two species became isolated from the Indo-Pacific by the Langhian eastern closure of the Mediterranean basin and disappeared during the Messinian Salinity Crisis. Since the Tortonian, B. schlumbergeri, which descended from B. philippinensis, has inhabited the Indo-Pacific along with B. pulchra. From the central Pacific Ocean, B. pulchra reached the Caribbean area before the early Piacenzian closure of the Central America seaway.