Two Species of Hesslandona (Phosphatocopida, Crustacea) from the Upper Cambrian of Western Hunan, South China and the Phylogeny of Phosphatocopida
Two Species of Hesslandona (Phosphatocopida, Crustacea) from the Upper Cambrian of Western Hunan, South China and the Phylogeny of Phosphatocopida
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DOI：
10.1666/10-043.1
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发表时间：
2011-07
期刊：
影响因子：
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通讯作者：
Huaqiao Zhang;Xi-ping Dong;S. Xiao
中科院分区：
文献类型：
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作者：
Huaqiao Zhang;Xi-ping Dong;S. Xiao
本文描述了在湖南西部上寒武统中发现的一个新的类磷化石Hesslandona necopina Müller，1964和Hesslandona longispinosa（Kozur，1974）。在个体发育阶段，H。necopina修订，新定义的第二和第三个体发育阶段。第二阶段的特征是下颌肢干由分开的基节和基足组成。第三阶段的特征是下颌基节和基足部分融合。这两个部分可能在稍后的个体发育阶段完全融合。长刺西虎鱼（Vestrogothia longispinosa Kozur），1974年被重新分配到Hesslandona，因为存在一个interdorsum和一个相对狭窄的doublure。本文结合2005年以来在湘西同一剖面和层位上的新资料，对该科的系统发育进行了分析。定义了几个单系群的同源性（例如，广义甲壳纲、Labrophora、Eucrustacea、Phosphatocopida、Euphosphatocopida、Vestrogothiina、Hesslandonina和Dorsospinata）。目前的分析证实了早期的系统发育分析的结果，显示Phosphatocopida-Eucrustacea姐妹类群的关系，但不同于他们在支持Hesslandonidae的并系和Vestrogothiidae的单系。
Abstract The phosphatocopids Hesslandona necopina Müller, 1964 and Hesslandona longispinosa (Kozur, 1974) new combination, recovered from the Upper Cambrian in western Hunan, South China, are described. The ontogenetic stages of H. necopina are revised, with newly defined second and third ontogenetic stages. The second stage is characterized by a bipartite mandibular limb stem consisting of separate coxa and basipod. The third stage is characterized by the partial fusion of the mandibular coxa and basipod. These two parts may become completely fused possibly in a later ontogenetic stage. Vestrogothia longispinosa Kozur, 1974 is reassigned to Hesslandona because of the presence of an interdorsum and a relatively narrow doublure. The new data reported here, plus earlier reports of phosphatocopids from the same section and horizon in western Hunan since 2005, are included in an updated phylogenetic analysis of the Phosphatocopida. Autapomorphies that define several monophyletic groups (e.g., the Crustacea sensu lato, Labrophora, Eucrustacea, Phosphatocopida, Euphosphatocopida, Vestrogothiina, Hesslandonina, and Dorsospinata) are discussed. The present analysis confirms results from earlier phylogenetic analyses in showing the Phosphatocopida-Eucrustacea sister-taxon relationship, but differs from them in supporting the paraphyly of the Hesslandonidae and the monophyly of the Vestrogothiidae.