Tackling the "Protoblattoidea problem": Revision of Protoblattinopsis stubblefieldi (Dictyoptera; Late Carboniferous)
Tackling the "Protoblattoidea problem": Revision of Protoblattinopsis stubblefieldi (Dictyoptera; Late Carboniferous)
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解决“Protoblattoidea问题”：Protoblattinopsis stubblefieldi（网翅目；晚石炭世）的修订
DOI：
10.14411/eje.2009.017
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发表时间：
2009
影响因子：
5.8
通讯作者：
J. Schneider
中科院分区：
文献类型：
--
作者：
O. Béthoux;K. Klass;J. Schneider
化石物种 †Protoblattinopsis Stubblefieldi Laurentiaux, 1953（网翅目；晚石炭世）的前翅形态被重新研究。与之前所述不同的是，在该物种中，R 和 M 的茎是不同的，而 CuA 有分支。 †P 中出现假定的类似“长 ScP”和“分化的 R 扇区”（即 RA 和 RP）。 Stubblefieldi 表示所有现存网翅目及其并系茎的许多成员的基部位置。与 †Archimylacridae 和 †Blattinopsidae 科的代表不同，但与 Blattodea s 相似。 str.，†P。 Stubblefieldi 缺乏 M-CuA 弓。 †P 中中位系统分支模式的比较。 Stubblefieldi 和 †Archimylacridae 表明后者的弓由 M 的后分支组成。值得注意的是，CuA 和 CuP 起源点的远端位置、CuA 前茎的乙状结肠走行、CuP 的强弯曲以及 R/RP 和 M 前分支之间短融合的发生与 Polyphaga Brulle 属的代表相似，1835 年，属于网翅目 s.海峡然而，这些相似的性状状态在这些类群中不太可能是同源的，主要是因为多食目是其余网翅目的姐妹类群的假设。海峡（甚至 Blattodea s. str.）没有得到强烈支持。由于其特征状态组合，该物种 †P。 Stubblefieldi 在“原蠼总科垃圾桶”中是独一无二的。需要对干网翅目的翅膀形态进行全面研究，以阐明它是否可能与网翅目的谱系密切相关。海峡进化了。
The forewing morphology of the fossil species †Protoblattinopsis stubblefieldi Laurentiaux, 1953 (Dictyoptera; Late Carboniferous) is re-investigated. Unlike previously stated, in this species, the stems of R and M are distinct and CuA has branches. The occurrence of the putative plesiomorphies "long ScP" and "differentiated R sectors" (i.e., RA and RP) in †P. stubblefieldi indicates a position basal to all extant Dictyoptera and many members of its paraphyletic stem. Unlike representatives of the families †Archimylacridae and †Blattinopsidae, but similar to Blattodea s. str., †P. stubblefieldi lacks a M-CuA arculus. The comparison of the branching pattern of the median system in †P. stubblefieldi and †Archimylacridae suggests that the arculus of the latter is composed of posterior branches of M. It is noted that the distal location of the point of origin of CuA and CuP, the sigmoid course of the anterior stem of CuA, the strong curvature of CuP, and the occurrence of a short fusion between R/RP and the anterior branch of M are similarities with the representatives of the genus Polyphaga Brulle, 1835, belonging to Dictyoptera s. str. However, these similar character states are unlikely to be homologous in these taxa, mainly because the hypothesis that Polyphaga is the sister group of the remaining Dictyoptera s. str. (or even Blattodea s. str.) is not strongly supported. Owing to its character state combination, the species †P. stubblefieldi is unique within the "Protoblattoidea waste-basket". Comprehensive studies of wing morphology in stem-Dictyoptera are needed to clarify whether it might be closely related to the lineage from which Dictyoptera s. str. evolved.