THE SUBTRIBES AND GENERA OF THE TRIBE BROSCINI (COLEOPTERA: CARABIDAE): CLADISTIC ANALYSIS, TAXONOMIC TREATMENT, AND BIOGEOGRAPHICAL CONSIDERATIONS
THE SUBTRIBES AND GENERA OF THE TRIBE BROSCINI (COLEOPTERA: CARABIDAE): CLADISTIC ANALYSIS, TAXONOMIC TREATMENT, AND BIOGEOGRAPHICAL CONSIDERATIONS
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布鲁西尼部落的亚属和属（鞘翅目：步甲科）：分类分析、分类学处理和生物地理学考虑因素
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发表时间：
2000
期刊：
影响因子：
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通讯作者：
S. Roig‐Juñent
中科院分区：
文献类型：
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作者：
S. Roig‐Juñent
摘要采用分支系统学的方法，对步甲科步甲族各属的系统发育关系进行了研究。共观察到88种昆虫的73个形态特征，隶属于41个属和亚属，并以Melaenini族和Apotomini族为外类群。在所有获得的分支图中，属被分为五个分支，作为新的亚族分类的基础：东方-古北区-新热带区Axonyina，新亚族（模式属，Axonya Andrewes）有3属;全北极-东方的Broscina亚族（模式属Broscus Panzer）9属;南温带澳大利亚-新热带区Nothobroscina新亚族（模式属Nothobroscus Roig-Juñent and Ball）有10个属;南温带新热带的Barypina（模式属Barypus Dejean）有2属;和南温带澳大利亚-新热带Creobiina（模式属Creobius Guérin-Ménéville）有11属。在不同的属Barypus，Eurylychnus Bates，和Promecoderus Dejean物种组被视为终端单位，以测试这些属的单系性。分析结果表明，只有Promecoderus不是单系的，目前的结构。提供了Broscini属的检索表和特征，并说明了所有分类学上重要的结构。雄性和雌性的生殖器特征是首次描述的几个属。本文报道了微蜂属Microbarypus的模式种M.新发现的森林（模式产地Alto Queulat，Aisén，智利）。同色小鲈Percosoma concolor Sloane、副纹状体小鲈P. substriatum摩尔和高山小鲈P. montanum Castelnau被转移到乳糜属。模式种为（学名：tela concolor Castelnau）、细颚（学名：Gnathoxys granularis Westwood）和蜡（学名：Cerotalis substriata Castelnau）。讨论了解释Broscini分布的四个主要地理学假说。最近新发现的新热带属，如Nothobroscus Roig-Juñent and Ball和Rawlinsius Davidson and Ball，以及Broscini的新属，导致了一个不同的地理假说。Broscini可以被认为是一个起源于下侏罗纪时代的泛古陆部落，当时南美洲和北美洲在太平洋边缘有陆地连接。从广义上讲，布罗斯奇尼的生物地理模式可以用目前盘古大陆解体的假说来解释。
Abstract Phylogenetic relationships of the genera of the tribe Broscini (Carabidae) are postulated based on cladistic methods. Seventy-three morphological characters were examined in 88 species, arranged in 41 genera and subgenera, and with the related tribes Melaenini and Apotomini as outgroups. In all the obtained cladograms, the genera are grouped in five clades recognized as the basis for a new subtribal classification: the Oriental-Palearctic-Neotropical Axonyina, new subtribe (type genus, Axonya Andrewes) with 3 genera; the Holarctic-Oriental subtribe Broscina (type genus Broscus Panzer) with 9 genera; the south temperate Australian-Neotropical Nothobroscina new subtribe (type genus Nothobroscus Roig-Juñent and Ball) with 10 genera; the south temperate Neotropical Barypina (type genus Barypus Dejean) with 2 genera; and the south temperate Australian-Neotropical Creobiina (type genus Creobius Guérin-Ménéville) with 11 genera. Species groups within diverse genera Barypus, Eurylychnus Bates, and Promecoderus Dejean were treated as terminal units to test monophyly of these genera. Results of the analysis show that only Promecoderus is not monophyletic, as presently structured. Keys and diagnoses are provided for the genera of Broscini, and all taxonomically important structures are illustrated. Genital features of males and females are described for the first time for several genera. The barypine genus Microbarypus, with its type species M. silvicola (type locality Alto Queulat, Aisén, Chile) is described as new. The species Percosoma concolor Sloane, P. substriatum Moore, and P. montanum Castelnau are transferred to the genus Chylnus. Type species are designated for Adotela Castelnau (Adotela concolor Castelnau), Gnathoxys Westwood (Gnathoxys granularis Westwood), and Cerotalis Castelnau (Cerotalis substriata Castelnau). The four main biogeographical hypotheses proposed to explain Broscini distribution are discussed. Recent discovery of new Neotropical genera, as Nothobroscus Roig-Juñent and Ball and Rawlinsius Davidson and Ball, together with the phylogeny of Broscini, lead to a different biogeographic hypothesis. Broscini can be considered a Pangaeic tribe originating in lower Jurassic times when there was a land connection between South America and North America on the Pacific margin. In broad terms, the biogeographic pattern of Broscini may be explained by the current hypothesis of the breakup of Pangaea.