Environmental Heterogeneity, Local Adaptation, and Sympatric Speciation in the Hawaiian Tree, Metrosideros Polymorpha

夏威夷树 Metrosideros Polymorpha 的环境异质性、局部适应和同域物种形成

• 批准号: 0542350
• 负责人: Elizabeth Stacy
• 金额: $ 17.4万
• 依托单位: University of Hawaii at Hilo
• 依托单位国家: 美国
• 项目类别: Standard Grant
• 财政年份: 2006
• 资助国家: 美国
• 起止时间: 2006-05-01 至 2008-10-31
• 项目状态: 已结题
• 来源: https://www.nsf.gov/awardsearch/showAward?AWD_ID=0542350&HistoricalAwards=false
• 关键词: Environmental; Heterogeneity; Local; Adaptation; Sympatric

Title: Environmental heterogeneity, local adaptation, and sympatric speciation in the Hawaiian tree, Metrosideros polymorphaAbstractSpecies in the early stages of diversification serve as excellent models for studies of the environmental conditions that drive speciation. The highly variable Hawaiian tree, Metrosideros polymorpha ('Ohi'a), is such a species, having derived from a progenitor that colonized the Hawaiian Islands from New Zealand as recently as 0.5 to 1.0 million years ago. Several of 'Ohi'a's varieties co-occur across the volcanic islands, which range in age from the environmentally stable, 5-million-year-old Kauai to the still-forming and highly heterogeneous Big Island. As such, 'Ohi'a provides an ideal system in which to examine the relationship between spatial and temporal environmental heterogeneity and the evolution of reproductive isolation, a precursor to speciation. This two-year study will use hand-pollinations, microscopy, and growth chambers to determine the relative cross-fertility within and between varieties and the viability of the resulting seedlings under a range of conditions. The proposed work will help to illuminate the environmental conditions under which speciation in out crossed tree species will occur. The work will involve the training of several undergraduate students and one M.S. student in evolutionary theory, field methods, and the scientific process. Recruitment of students will target under-represented groups at UH-Hilo, a Minority and Native Hawaiian-serving Institution. Further, the research projects developed through this grant will be modified by the P.I. into lecture and laboratory exercises in upper-undergraduate and graduate courses.

环境异质性、局部适应和同域物种形成 夏威夷树，Metrosideros polymorphaAbstract物种在多样化的早期阶段作为一个很好的模式，研究环境条件，驱动物种形成。 高度变异的夏威夷树Metrosideros polymorpha（'Ohi'a）就是这样一个物种，它的祖先在50万到100万年前从新西兰殖民到夏威夷群岛。 几个'Ohi' a '的品种共同出现在火山岛，其年龄范围从环境稳定，500万年的考艾岛到仍在形成和高度异质的大岛。 因此，“Ohi 'a”提供了一个理想的系统，在其中检查空间和时间的环境异质性和生殖隔离的演变，物种形成的前兆之间的关系。 这项为期两年的研究将使用人工授粉，显微镜和生长室来确定品种内和品种间的相对杂交生育力以及在一系列条件下产生的幼苗的生存能力。 拟议的工作将有助于阐明环境条件下，物种形成的外杂交树种将发生。 这项工作将涉及几名本科生和一名硕士的培训。学生在进化论，领域的方法，和科学的过程。 学生的招聘将针对UH-Hilo的代表性不足的群体，UH-Hilo是一个少数民族和夏威夷土著服务机构。 此外，通过这笔赠款开发的研究项目将由P.I.在高年级本科生和研究生课程的讲座和实验练习中。