PALYNOLOGY AND EMBRYOLOGY IN EUPHORBIACEAE

用扫描电镜（SEM）和透射电镜（TEM）分析了桉树科18个部落中17个部落64属110种的花粉（sensu Webster, 1989）。这个亚科在孢粉学上是多样的。cheilose科（Cheilosa）有棘状三羽花粉。在染色蕨科中，双轴属植物具有独特的双侧花粉，具有两个带孔。在小球果科，小球果有一个完整的顶盖，小球果呈网状，小球果呈细网状或微网状。阿德利亚科和阿尔乔尼亚科共享带孔的三聚花粉。桉树属的花粉粒具有异质性，具有不同的外壁雕刻和结构。棘科的特征是网状的外壁。Pereae有不寻常的由相互交织的小柱组成的外壁结构。大多数桉树科有连体孔，但孔的数量各不相同。广泛的花粉多样性表明，粉螨亚科是一个包括多个异质类群的多系分类单元。

Pollen of 110 species representing 64 genera from 17 of the 18 tribes in the Acalyphoideae (sensu Webster, 1989) was examined in SEM and TEM.This subfamily is palynologically diverse. The Cheiloseae (Cheilosa) have spinulate tricolporate pollen. In the Chrozophoreae, Ditaxis has unique bilateral pollen with two zonapertures. In the Pycnocomeae, Pycnocomas has a complete tectum with spinules in a reticulate pattern and Argomuellera is finely or reticulate or microreticulate. The Adelieae and the Alchornieae share operculate tricolporate pollen. The Acalypheae have heterogenous pollen grains with various exine sculptures and structures. The Dalechampieae are characterized by reticulate exines. The Pereae have unusual exine structure composed of interwoven columellae. Most Acalyphoideae have colporate apertures, but the number of apertures varies. The wide range of pollen diversity suggests that the subfamily Acalypohoideae is a polyphyletic taxon including several heterogeneous groups.