Studies on the tuberculous, mycorrhizic gametophytes and their embryogenesis in pteridophytes.

本研究的目的是获得更多的信息，结核性，菌根配子体，并澄清配子体功能的多样性。对裸盖蕨、石松和多裂阴地蕨的配子体和胚进行了观察。从形态学上描述了它们的特征，并对裸盖蕨科（Psilotaceae）、石松科（Lycopodiaceae）和瓶舌科（Ophioglossaceae）的结核性配子体进行了历史回顾。配子体纤毛状，基部倒圆锥形，几乎都有不等的二歧分枝。从自然界采集到的棒石松配子体约80个，包括从幼原叶体到大型成熟配子体的各种形态，并对其外部和内部形态进行了研究。本文对多裂阴地蕨（Botrychiummultifidum）的胚的解剖学特征进行了详细描述。成熟的精子为块茎状，背腹稍扁平，具一条中间的纵脊，其上产生精子器。年轻的胚胎保持一个广泛的椭圆形轮廓。胚柄由数个细胞组成，稍弯曲，嵌在配子体组织内。

The present study was designed to obtain additional information concerning the tuberculous,mycorrhizic gametophytes and to clarify the diversity of the gametophyte features. The gametophytes and embryos were observed for Psilotum nudum, Lycopodium clavatum and Botrychium multifidum. Observations on the features of them were described morphologically, together with the historical reviews on the tuberculous gametophytes in Psilotaceae, Lycopodiaceae and Ophioglossaceae.About two hundred gametophyte of Psilotum nudum were collected from a clay pot in which the sporophyte was growing. The gametophytes are cilindrical with obconic basal tip and almostly have unequal dichotomous branches. They have a strong resemblance to the slender small rhizomes having the same rhizoids and gemmae.About 80 gametophytes of Lycopodium clavatum from nature, including various form from young prothallia to large mature ones, were collected and the external and internal morphology were investigated. Especially the anatomical feature of the embryos are particularly described in this report.About two hundred gametophytes of Botrychium multifidum were observed. The mature one is tuberous, somewhat flattened dorsiventrally, having a median longitudinal dorsal ridge upon which antheridia are produced. The young embryo maintained a broadly oval outline. The suspensor is composed of several cells, and is slightly curved, embedded within the tissue of gametophyte.