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Systematics of Aneuraceae (Hepaticae) based on differentiation of thallus

基于叶状体分化的动脉科（肝科）系统学

基本信息

批准号：
13640712
负责人：
FURUKI Tatsuwo
金额：
$ 1.73万
依托单位：
Natural History Museum and Institute, Chiba
依托单位国家：
日本
项目类别：
Grant-in-Aid for Scientific Research (C)
财政年份：
2001
资助国家：
日本
起止时间：
2001 至 2004
项目状态：
已结题

来源：
https://kaken.nii.ac.jp/grant/KAKENHI-PROJECT-13640712/
关键词：
Southeast Asia Bryophytes Hepaticae Aneuraceae Thallus Oilbody tissue differentiation Systematics 系統分類

项目摘要

1.Young stages developed from sporeling : Oil bodies are included in all cells of young stage derived from species having oil bodies in all cells in adult. On the other hand, species without oil bodies in epidermal or marginal cells in adult lack oil bodies in early young stages and became present in only inner cells, or early young stage includes oil bodies all cells and become to lack them gradually in epidermal cells.2.Differentiation of thalli based on oil bodies : The thalli have interdependence between tissue differentiation and oil body-distribution. (1)All cells include oil bodies : no defferentiation of thallus-Aneura, Lobatiriccardia and some species of Riccardia subg. Riccardia. (2)Oil bodies scattered or lacking-Riccardia subgen. Riccardia. (3)Ventral epidermal and marginal cells lack oil bodies-Riccardia subg. Hyaloneura. (4)Oil bodies only in inner cells : thallus has small cells-some species of Riccardia subg. Riccardia. (5)Inner and part of wing cells include oil bodies : having gaps between epidermal cells and wing-some species of Riccardia subg. Riccardia.3.Reexamination of type specimens : Thirty-nine species described from Southeast Asia of the family are later homonyms and must be reduced under other earlier correct name as synonyms.4.Taxonomical status of Vandiemenia : Thalli have no oil bodies and midrib structures and they suggest that it belongs to the family Metzgeriaceae.5.Systematics : Five species of Ancura, 2 species of Lobatiriccardia, 45 species of Riccardia are rccognized in Southeast Asia. Among them 1 species of Ancura, 1 species of Lobatiriccardia, 4 species of Riccardia may be described as new to science. One species of Tasmania may be also described as new to science. About ten species arc open to question and further investigations are needed.

1.从孢子发育而来的幼龄阶段：幼龄阶段的所有细胞中都含有油体，这些油体来自于成体所有细胞中都具有油体的物种。另一方面，成虫表皮或边缘细胞中没有油体的物种，在幼龄早期缺乏油体，只存在于内部细胞中，或者幼龄早期包括所有细胞的油体，但表皮细胞中逐渐缺乏油体。 2.基于油体的菌体分化：菌体的组织分化和油体分布之间存在相互依赖性。 (1)所有细胞均含油体：叶状体-Aneura、Lobatiriccardia和某些Riccardia subg种无分化。里卡迪亚。 (2)油体散在或缺失-Riccardia subgen。里卡迪亚。 (3)腹表皮和边缘细胞缺乏油体——Riccardia subg。透明质酸。 (4)油体仅存在于内细胞中：叶状体有小细胞——Riccardia subg.的一些种。里卡迪亚。 (5)翅细胞的内部和部分含有油体：表皮细胞与翅细胞之间有间隙-Riccardia subg的一些种。 Riccardia.3.模式标本的重新审查：来自东南亚的该科的 39 个物种是后来的同音异名，必须减少到其他早期正确名称下作为同义词。4.Vandiemenia 的分类地位：Thalli 没有油体和中脉结构，他们认为它属于 Metzgeriaceae 科。5.系统学：Ancura 的 5 种，Ancura 的 2 种 Lobatiriccardia，东南亚已发现 45 种 Riccardia。其中Ancura属1种、Lobatiriccardia属1种、Riccardia属4种堪称科学新物种。塔斯马尼亚的一种物种也可能被描述为科学上的新物种。大约有十个物种有待商榷，需要进一步调查。