Were huge ancient lakes refugia for fishes distributed in Mogami River System ?

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The recent geological data suggests that two huge lakes had developed around the upper and lower reaches of recent Mogami River System during 4 - 6 million years ago. We conducted population genetic analyses of seven fishes that might be evolved around the ancient lakes, and discussed the influence of ancient lakes for their evolution. Two of the seven species had been probably speciated around the ancient lakes. In other three species, genetic structures were different between Mogami and adjacent river populations or between the upper and lower reach populations in Mogami River System, suggesting that geographical isolation by the ancient lakes had affected to their genetic differentiation. In other two species, distinctive genetic structures did not recognize between populations. In Sagami River populations to compare with Mogami River populations, clear genetic analyses could not do by remarkable introduction of individuals derived from different river systems.