环境异质性对克隆植物基株内和基株间作用关系的影响

资源分布的时空异质性在自然界普遍存在，并深刻影响着各种生态学格局与过程，但目前国内外有关时空异质性对克隆植物基株内和基株间作用关系影响的研究仍十分缺乏。以入侵我国南方地区的外来克隆植物香菇草为研究对象，运用实验生态学、分子生态学和稳定性同位素等方法，研究植物必需资源（即光照、土壤养分和水分）的空间异质性和时间异质性对克隆植物不同基株的分株之间及相同基株的不同独立分株和相连分株之间作用关系的影响，检验假说“如果克隆植物不同基株在异质性环境中的觅食能力存在显著差异，那么资源分布的时空异质性可显著影响基株间的作用关系”、“资源分布的时空异质性不显著影响基株内独立分株之间的作用关系”和“资源分布的时空异质性对基株内相连分株之间作用关系的影响与分株之间的克隆整合能力密切相关”，揭示资源分布的时空异质性对克隆植物基株内和基株间作用关系的影响及机制，丰富和发展植物竞争理论和克隆植物生态学的理论与方法。

Both spatial and temporal heterogeneity in resource supply are common in nature, and can significantly affect various ecological patterns and processes. So far, however, very few studies have examined their effects on intra or inter-genet interactions of clonal plants. Taking Hydrocotyle vulgaris L., a introduced clonal herb that is invasive in southern China, as a mode plant and using the methods of experimental ecology, molecular ecology and stable isotope, we will investigate the effects of spatial and temporal heterogeneity in light, soil nutrient and water supply on the interactions between isolated ramets of different genets (genotypes) and between isolated or interconnected ramets of the same genets. We will test the hypotheses (1) that spatial and temporal heterogeneity in resource supply will significantly affect inter-genet interactions if the foraging ability of different genets differs, (2) that spatial or temporal heterogeneity in resource supply does not affect the interactions between isolated ramets of the same genets, and (3) that the effects of spatial and temporal heterogeneity in resource supply on the interactions between interconnected ramets of the same genets depend on the ability of clonal integration (resource sharing) between the interconnected ramets. The aim of this project is to disclose how spatial and temporal heterogeneity in resource supply affects intra- and inter-genet interactions of clonal plants in order to enrich the theories and methods of plant competition and clonal plant ecology.