二型花柱作为被子植物花性器官演化和物种形成过程中的主要问题已引起进化生态学领域的高度关注，但二型花柱植物的起源和演化十分不清楚。因此，不同花型近缘种繁育系统的比较研究，对揭示二型花柱物种起源和演化趋势具有重要的意义。雪地报春(Primula nivalis)和突厥报春(P. turkestanica)是在我国仅分布于新疆的不同型姐妹种；二者在生物学特性、空间分布特点及生长微环境间存在差异。因此，我认为二型花柱雪地报春迁移到新居群或繁殖过程中，与居群海拔有关生态因子变化导致的繁育系统演化，可能是形成次生同型花柱突厥报春的主要因素。为验证该假设，本项目将通过对雪地报春和突厥报春的花部特征分化和居群遗传结构、传粉特性以及交配系统的比较研究。揭示这两物种在繁育系统特点上的异同点；探讨这些特点在二者演化过程中的作用，以期为全面揭示二型花柱植物繁育系统演化趋势并进一步探讨被子植物花分化机制提供理论依据

Research about new species evolution process has fascinated hundreds of plant evolutionary ecologists. Although, distyly species is main group of angiosperm flower morph evolution the most of distyly species are still poorly understood for their origin, evolution, adaptation and maintenance. The comparative study on two different floral morph sister species breeding system plays an important role for investigation on origin and evolution in homostyly from distyly morph. Primula nivalis and P. turkestanica are two different floral morph sister species viz. homostyly and distyly of Primula, and grow only in northern Xinjiang, China. The biological characters, population characters, and growth area are different for these species in natural populations. Present study aimed to study on evolution of homostyly species from distyly and determination of the floral differenciation in sister species. We hypothesized that the evolution of homostyly P. turkestanica, when distally species of P. nivalis was migrated to a new population or breeding process and the changes in ecological factors (altitude or pollinator) associated to the evolution of P. nivalis breeding systems, were the major factor for speciation of secondary homostyly P. turkestanica. To test this hypothesis, the effects of ecological factors on differentiations of floral character, population genetic structures, pollination, and mating system will be quantified for two sister species of Primula in Xinjiang. At the completion of this study, we will ascertain the dynamic breeding system and evolution history of two different floral morph sister speceis. Moreover, we will explore the ecological adaptive strategies and effect of change of population structure dependent breeding system on evolution prosees of these two sister speceis. Our overall goal is to develop a general theoretical basis for evolution of distlly speceis and determination of the floral differenciation of angiosperm species. ..