中国东北中生代草蛉类化石系统分类与演化研究

草蛉类化石在我国东北中生代地层中较为常见，是燕辽和热河昆虫群的重要组成部分。全球已知最早的草蛉科化石即采自我国中侏罗统九龙山组，并在下白垩统义县组中大量出现，物种更为多样，它们是草蛉科起源与早期辐射演化的重要代表。此外，草蛉类昆虫的基干类群也在中生代大量出现。目前草蛉科系统发育关系争议较大，而草蛉类中的多个中生代绝灭科与其近缘现生科的系统发育关系尚不明确。本研究拟基于我国大量的化石标本，通过经典分类学研究，厘清中生代草蛉类昆虫多样性组成；在此基础上，结合化石与现生类群，整合形态学特征与DNA分子序列，构建全证据矩阵，进行系统发育分析；并重建重要形态特征的祖先状态和演化历史，估算各分支节点的分歧时间；旨在获得完整地质历史的草蛉科系统发育树，追溯中生代草蛉类昆虫的辐射演化历史。研究结果将为脉翅目的系统演化研究和重建燕辽、热河昆虫群提供重要线索和直接证据。

Fossil chrysopid-like lacewings have been extensively collected in the late Mesozoic of northeastern China, forming important components of Yanliao Biota and Jehol Biota. The Middle Jurassic Jiulongshan Formation yielded the earliest fossil records of Chrysopidae over the world. Consequently, more diverse chrysopids were found in the Lower Cretaceous Yixian Formation. In addition, stem groups of the chrysopid-like lacewings bloomed in the Mesozoic. They are essential for studying the origin and early evolutionary radiation of the clade. To date, the phylogeny of Chrysopidae is still controversial, as well as the clade of chrysopid-like lacewings. Phylogenetic relationships among the extinct families of Mesozoic and their closely related extant families are unclear. In this proposal, a comprehensive investigation of chrysopid-like lacewings will be performed, based on abundant fossil specimens from the Mesozoic of China. Morphology and taxonomy will be studied and reviewed though their entire evolutionary history. Total-evidence phylogeny will be analyzed integrating fossil and extant species, using morphological characters and DNA sequences. Venation transformation and their ancestral states will be reconstructed. Divergence estimates will be conducted for nodes on the chrysopid-like lacewings tree. The results of this proposed study will be valuable references for future studies of systematics and evolution of Neuroptera and reconstruction of Yanliao Biota and Jehol Biota.