薹草属细柄薹草组(莎草科)的系统分类研究
批准号:
32070218
项目类别:
面上项目
资助金额:
58.0 万元
负责人:
金孝锋
依托单位:
学科分类:
植物分类学
结题年份:
2024
批准年份:
2020
项目状态:
已结题
项目参与者:
金孝锋
中文摘要
薹草属Carex是被子植物的大属,全世界1960~2000种。中国有520余种,分布广泛,生境多样,该属的分类历来被认为是一大难题。随着深入的调查采集和植物志编撰,薹草属的分类学和系统学研究均取得了较大进展,但在种的划分、属内组的界定等方面仍存在问题和严重分歧,反映出以往对薹草属分类研究方法单一,对多学科综合评价和分析系统分类问题的认识不足。.本研究坚持居群概念,对主产东亚的细柄薹草组sect. Graciles植物进行广泛深入的野外调查和标本采集,馆藏标本的全面查阅和观测,用统计学方法分析种间及种内居群间的形态变异,探讨和评价形态性状(包括宏观形态,叶表皮和解剖结构、果囊及小坚果的微形态)的变异式样及其系统分类学意义,结合转录组数据,揭示本组的系统发育关系、进化起源中心和分化时间,并进行全面系统分类研究。本项目既为Carex系统分类研究提供借鉴,也培养植经典分类和系统进化研究的后备人才。
英文摘要
Cyperaceae, a monocot family in the order Poales, contains 106 genera and ca. 5400 species worldwide, with closest relatives, Juncaceae and Thurniaceae. With the amount of 1960 to 2000 species, Carex L. is the largest genus in the family Cyperaceae. The genus has traditionally been divided into four subgenera: subg. Vigneastra (Tuck.) Kük., subg. Vignea (B. Beauv. ex Lestib.) Peterm., subg. Psyllophora (Degl.) Peterm., and subg. Carex. More recently, these subgenera are known to be largely polyphyletic or paraphyletic, with the notable exception of subg. Vignea. The genus Carex itself, however, is cosmopolitan and easily distinguished from the other genera of Cyperaceae based on the combination of unisexual flowers and achenes enveloped in perigynia..With a number of species and a large dispersal region, and limited specimens, the taxonomic revision and floristic studies on Carex are notorious and considered as one of the difficult problems in angiosperm. During the preparation of the regional florae, the investigation and taxonomy of Carex were fairly in-depth in the recent 40 years, but a lot of divergences and problems in species delimitation, generic subdivision, and geographical distribution etc. are still awaiting to solve. All these problems indicate that the cognition of morphological characters and its systematic significance are far from a satisfactory status..Based on field work, population sampling, specimen examination of the main herbaria both at home and abroad, this project will use statistical analysis and analyze the variation of the morphological characters, which are used to identify the species in Carex sect. Graciles. We will also evaluate the taxonomic significance/value of these characters, including micormorphology of nutlets, perigynia, anatomy and epidermal characters of leaves, conduct phylogenetic analysis of transcriptomic data on all sectional species, as well as some problematic species (Carex brunnea and C. gentilis) with their allies, and make a full-scale revision of Carex sect. Graciles, as well as delimitate this section and similar ones. It can accelerate the taxonomic revision, systematics, origin, dispersal and the floristic studies of the whole genus Carex. The project will benefit for cultivating reserve talents of phytotaxonomy.
薹草属细柄薹草组Carex sect. Graciles是薹草亚属近基部的一支,主产东亚,分类学问题突出。本研究在标本查阅、野外采集、标本鉴定的基础上,对本组进行了果囊和小坚果扫描电镜观察、叶解剖和叶表皮实验、基于二代测序的系统发育分析和对复合体的形态性状变异研究,并结合这些证据最终对细柄薹草组进行界定和分类修订,主要结果如下:(1)果囊的形态和微形态在种内表现出稳定性,果囊颜色和长度可用于界定细柄薹草组,具有系统学意义;果囊的形状、表面毛被、喙长、喙口形态在区分物种时有价值。(2)根据小坚果的形态和微形态显示种内稳定性,在组内也具较高的一致性,小坚果长度在界定细柄薹草组时有价值。(3)叶片解剖结构在组内较为一致,可作为区分某些种的依据。(4)叶表皮特征在大多数种间表现出稳定性。(5)基于片段的系统发育分析分辨率和支持率不高,但在组的界定中有价值;基于基因组重测序的系统发育研究进一步明确了一些种的种级地位,对该组的起源历史有一定启示。(6)形态性状变异研究为复合体内种和种下等级的界定提供了依据。(7)修订后承认了本组全世界共19种,发现2新种,1新组合变种,7个新异名。(8)长穗柄薹草C. longipes、圆坚果薹草C. orbicularinucis和华疏花薹草C. sinodissitiflora等不属于细柄薹草组。
薹草属细柄薹草组(莎草科)的系统分类研究
-
批准号:--
-
项目类别:--
-
资助金额:58万元
-
批准年份:2020
-
负责人:金孝锋
-
依托单位:
东亚-北美间断分布稀花薹草组Carex sect. Paniceae(莎草科)的系统分类研究
-
批准号:31770212
-
项目类别:面上项目
-
资助金额:57.0万元
-
批准年份:2017
-
负责人:金孝锋
-
依托单位:
世界薹草属密花薹草组(莎草科)的分类修订
-
批准号:31470304
-
项目类别:面上项目
-
资助金额:80.0万元
-
批准年份:2014
-
负责人:金孝锋
-
依托单位:
薹草属广义灰帽薹草组(莎草科)的系统分类研究
-
批准号:31170183
-
项目类别:面上项目
-
资助金额:55.0万元
-
批准年份:2011
-
负责人:金孝锋
-
依托单位:
薹草属菱形果薹草组(莎草科)的分类研究
-
批准号:30870150
-
项目类别:面上项目
-
资助金额:25.0万元
-
批准年份:2008
-
负责人:金孝锋
-
依托单位:
国内基金
海外基金