Systematics of Aneuraceae (Hepaticae) based on differentiation of thallus

1.由散裂发育而来的幼期：由成虫的所有细胞中都有油体的物种衍生而来的幼期的所有细胞中都含有油体。另一方面，成虫表皮或边缘细胞中没有油体的种在幼嫩期早期缺乏油体，只在内部细胞中存在，或幼嫩期早期包括所有细胞的油体，逐渐在表皮细胞中缺乏油体。基于油体的菌体分化：菌体的组织分化与油体分布之间存在着相互依存关系。(1)所有细胞均含油体：未分化成瘤状瘤、叶状瘤和部分瘤状瘤亚种。Riccardia。(2)油体分散或缺油；Riccardia。(3)腹侧表皮和边缘细胞缺乏油体- riccardia亚群。Hyaloneura。(4)油体只存在于内细胞：菌体中有小细胞——某些种类的Riccardia亚属。Riccardia。(5)翅细胞内部及部分含油体，表皮细胞与翅之间有间隙。Riccardia.3。模式标本的复验：在东南亚所描述的该科中有39种是后来的同音异义，必须归入其他较早的正确名称作为同义词。4 . Vandiemenia菌体的分类地位：菌体无油体，无中脉结构，属于metzgeriaceae科。分类学：在东南亚地区已发现的安科属5种，罗氏属2种，李氏属45种。其中安科属1种，罗氏属1种，李氏属4种为新属。塔斯马尼亚岛的一个物种也可能被描述为科学上的新物种。大约有10种尚待研究，需要进一步调查。

1.Young stages developed from sporeling : Oil bodies are included in all cells of young stage derived from species having oil bodies in all cells in adult. On the other hand, species without oil bodies in epidermal or marginal cells in adult lack oil bodies in early young stages and became present in only inner cells, or early young stage includes oil bodies all cells and become to lack them gradually in epidermal cells.2.Differentiation of thalli based on oil bodies : The thalli have interdependence between tissue differentiation and oil body-distribution. (1)All cells include oil bodies : no defferentiation of thallus-Aneura, Lobatiriccardia and some species of Riccardia subg. Riccardia. (2)Oil bodies scattered or lacking-Riccardia subgen. Riccardia. (3)Ventral epidermal and marginal cells lack oil bodies-Riccardia subg. Hyaloneura. (4)Oil bodies only in inner cells : thallus has small cells-some species of Riccardia subg. Riccardia. (5)Inner and part of wing cells include oil bodies : having gaps between epidermal cells and wing-some species of Riccardia subg. Riccardia.3.Reexamination of type specimens : Thirty-nine species described from Southeast Asia of the family are later homonyms and must be reduced under other earlier correct name as synonyms.4.Taxonomical status of Vandiemenia : Thalli have no oil bodies and midrib structures and they suggest that it belongs to the family Metzgeriaceae.5.Systematics : Five species of Ancura, 2 species of Lobatiriccardia, 45 species of Riccardia are rccognized in Southeast Asia. Among them 1 species of Ancura, 1 species of Lobatiriccardia, 4 species of Riccardia may be described as new to science. One species of Tasmania may be also described as new to science. About ten species arc open to question and further investigations are needed.