Mutimodal learning in butterflies

这项研究考察了蝴蝶在觅食花朵的过程中是否能够学习两种模式，即颜色和气味。第一个实验表明，菜粉蝶可以通过奖惩双重条件反射来学习花香。第二个实验表明，它们也可以通过单独的奖赏条件反射来学习花香。第三个实验表明，蝴蝶不仅可以同时学习花卉的颜色和气味，而且它们可以通过学习两个通道而不是一个通道来提高辨别花朵的能力。

This study examined whether butterflies can learn two modalities, i.e., color and odor, during foraging for flowers. The first experiment revealed that Peris rapae butterflies can learn floral odor by dual conditioning of reward and punishment. The second experiment revealed that they also can learn floral odor by reward conditioning alone. The third experiment revealed not only that butterflies can learn floral color and odor simultaneously, but also that they can enhance the abilities to discriminate flowers by learning two modalities rather than one modality.