Clarification of key enzyme for ammonium assimilation under excess condition of ammonium to develop a tolerated cultivar against ammonium stress

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To get the basic information to give some leafy vegetables, which like nitrate nitrogen, tolerance of ammonium nitrogen, I examined effects of excess ammonium stress on growth and ammonium assimilation in head lettuce which can tolerate it and spinach mustard which is susceptible to it. The results suggest that the key enzyme of ammonium assimilation in both leafy vegetables under excess ammonium stress was glutamine synthetase(GS). But when GS was deactivated by serious ammonium stress, glutamate dehydrogenase(GDH) could mainly assimilate ammonium in the stead of Gs.