Spatial determinants of food web structure and function, from landscape to global scales

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How do food webs change over space? One the one hand, distant food webs may contain different species because of differences in the local environment or limited dispersal. Such differences in food webs may be extreme when they are so far apart as to draw from different species pools. On the other hand, even though species composition differs, the traits of the species may remain the same, such that food webs “work” the same way. Resolving the effects of space on the structure and functioning of food webs is critical for understanding how large-scale stressors (e.g. climate change, forest fragmentation) affect food webs. To resolve how space affects food webs, we need to (a) replicate experiments over landscape and continental scales, to determine how dispersal or biogeography changes species or traits; (b) expose such replicated food webs to environmental change, like altered climate, and reduced dispersal, such as occurs in isolated forest fragments; and (c) transplant species or even food webs between sites, to separate site-differences in species traits from the environment.