具粗细根茎植物的克隆断裂对环境异质性的适应对策

同时具直立粗根茎和横走细根茎的多年生草本是克隆植物中重要类型，粗细根茎分工使该类物种具有克隆断裂与克隆片段扩展的特殊性。本研究以具粗细根茎的克隆植物鸢尾属蝴蝶花为材料，通过以光、水与养分资源构成的多资源空间异质性（包括资源一致斑块性和资源交互斑块性等）处理的温室控制实验，结合同位素示踪、分子生态学实验与野外跟踪实验，分析异质性环境中该物种细根茎特征与断裂、细根茎断裂前克隆整合对植株生存与生长的影响，细根茎断裂后不同实验种群和野外自然种群克隆片段的生存与生长、种群动态与克隆结构动态，及片段扩展，以进一步探讨该物种细根茎特征（细根茎断裂前）、细根茎断裂现象、克隆整合与细根茎断裂早晚的权衡，及克隆片段的生存与扩展（细根茎断裂后）对环境异质性的适应与响应，最终阐明具粗细根茎植物的克隆断裂（及断裂后克隆片段扩展）对环境异质性的适应对策，为根茎克隆植物对异质性环境适应对策研究提供理论基础与科学依据。

Perennial herb with fine and coarse rhizome is one of significant types in clonal plants, which are possessed of specific characteristics in rhizome fragmentation and expansion of clonal fragments due to division of labor in rhizomes. Traits and fragmentation of fine rhizome, effect of clonal integration on the survival and growth before clonal fragmentation, the survival and growth, dynamics of ramet population and clonal structure, and expansion of clonal fragments in experimental populations and natural populations after clonal fragmentation in a clonal plant, Iris japonica with fine and coarse rhizome, were analyzed by greenhouse experiments with spatial heterogeneity (reciprocal patchiness and coincident patchiness etc.) of multiple resources, light, water content and nutrient in soil, combined with isotope tracers, molecular ecology experiments and field tracking experiments. Further, we will discuss the adaptation and response of traits and fragmentation of fine rhizome, trade-off between clonal integration and time for rhizome fragmentation, survival and expansion of fragments to environmental heterogeneity. Thus, adaptive strategies of clonal fragmentation (with fragment expansion) of plant with coarse and fine rhizome to environmental heterogeneity will be understood, which will provide a theoretical data and basis for adaptive strategies of rhizomatous clonal plants to environmental heterogeneity.