蓖麻是世界上重要的油料作物，尚未实现三系配套育种，花序发育分子机制是其研究的热点与难点。课题组以具有自主知识产权的蓖麻育种母本材料——带有标志性状的Lm型雌性系后代为研究对象，在前期工作基础上——发现PIP5K家族6个基因与Lm型雌性系后代花序发育直接相关，提出假说：PIP5K家族6个基因在蓖麻Lm型雌性系后代花序发育中存在功能，协同/递补调控其发育出单雌、标雌、两性系三种花序类型。本课题拟采用生物信息学、同源过表达和敲减表达等技术，明确PIP5K家族6个基因在花序发育过程中的功能；采用转录组学和蛋白质组学等手段，明确6个基因过表达和敲减表达时有哪些基因和蛋白质发生改变及其参与的代谢通路，6个基因间如何通过协同/递补作用完成对花序发育的调控，揭示PIP5K基因家族调控蓖麻Lm型雌性系后代花序发育的分子机制，也为其他类型蓖麻花序发育分子机制研究奠定基础。

Castor (Ricinus communis L.) is one of the most important oilseed crops in the world. However, three-line breeding has not been achieved in castor. In this study, we used Lm type female line progeny of castor as material to analyze the molecular mechanism of inflorescence that is a hot and difficult point. The research group has castor plant breeding parent material and it has independent intellectual property rights. Therefore, the study group is the research object of the Lm type female line with marked trait progeny of castor. Based on the previous studies, we found that six genes of PIP5K family involved in the development of inflorescence in Lm type female line progeny of castor and related to regulation of three types of inflorescence including to isofemale lines, markered female lines and monoecious lines. Here, we detected the six gene functions in the process of inflorescence development in Lm type female line progeny of castor by bioinformatics, gene overexpression and knock-down technologies. Then we also analyzed the related genes and proteins using transcriptomics and proteomics to study its function in the metabolic pathways. And how six genes regulated inflorescence development by synergistic/complementary effects. This study will reveal that the molecular mechanism on regulation of the PIP5K family to the development of inflorescence in Lm type female line progeny of castor, lay the foundation for the study of the molecular mechanism on other types of castor inflorescence development.