文化传扬演进与历史空间存续更新是历史城镇可持续发展的核心。我国历史城镇保护与更新改造的现实发展割裂了多元空间与文化生态的动态有机联系，导致城镇空间冻结在某一历史时刻或破坏传统文化生存的根基。如何促使二者协同共生，实现保护与更新的统一，历史性与现代性的统一，发掘其协同适应的内在机理并建立有效的协同机制是解决此类问题的关键。为此，本课题采用多学科交叉梳理与典型案例剖析、系统分析与系统综合、聚类归纳与对比研究、数据分析与动态模拟相结合等方法，发掘历史城镇空间与文化生态协同适应的本质关联要素及系统构成，辨识协同适应的层次类型及质性特征，揭示协同适应的动态过程及路径机制，并由此形成历史城镇空间与文化生态协同发展的多维解析框架。这既是对文化生态学视角下研究城市空间生态化建设理论的拓展，也是对历史文化遗产保护协同发展思想的深化与完善，更为历史城镇渐进式更新实践提供一种新思路与新解析框架。

Both the dissemination and evolution of culture and the existence and renewal of historical space are the key issues of sustainable development about historical towns. In fact, the protection and renewal of historical towns has separated the dynamic organic connection between multivariate space and cultural ecology, which has made the space of historical towns frozen in a particular historical moment or destroyed the foundation of traditional culture to survive in China. In order to promote the collaborative symbiosis between them, and realize the unification of protection and renovation, the unification of historic and modernity, it is critical to discover its inner mechanism of collaborative adaptation and establish an effective coordination mechanism. Therefore, this topic explores the essential associated elements and system structure of coordinated adaptation about historical towns space and cultural ecology, identifies their levels and types and qualitative characterization of coordinated adaptation, reveals the dynamic process and path mechanism about coordinated adaptation, and thus evolves into a multidimensional analytical framework about space of historical towns and cultural ecology with methods as multidisciplinary cross-combing and case analysis，systematic analysis and synthesis ,cluster induction and comparative study, data analysis and dynamic simulation. It is not only an expanding about the research on ecological construction theory of urban space, but also a deepening and improvement about collaborative development concept of historical and cultural heritage protection, which provides a new idea and a new analytical framework for the gradual renewal practice about historical towns.