植物适应光强波动变化的能力是决定光合产物累积的关键因素，健全的光破坏防御机制是动态光下光合机构有效运转的必要保障。研究发现UV-B可提升光破坏防御能力，但UV-B介导的光破坏防御机制形成过程还不清楚。本项目以番茄种苗为对象，拟通过UV-B处理诱导光破坏防御机制形成，从而提升种苗移植后对动态光的适应能力。首先，利用植物功能结构模型及解剖学手段，研究动态光下UV-B调控的植株表型和叶绿体运动对光截获能力的影响，阐明植物光捕获机制对UV-B的响应机理。其次，基于叶绿素a荧光动力学方法研究UV-B介导的光合机构能量耗散过程对动态光的响应；利用组学技术研究UV-B介导能量耗散过程与UVR8光受体及其下游信号伴侣的网络关系和关键基因。最后，研究UV-B介导光破坏防御机制对设施栽培条件下种苗光合性能的影响。本研究将有助于深入了解UV-B影响光合作用的生理本质，以期为设施种苗壮苗生产提供理论依据。

Plants’ capacity of acclimation under dynamic light plays a crucial role for photosynthates accumulation in nature, and a sound photoprotection mechanism is indispensable to guarantee effective operation of photosynthetic apparatus under dynamic light. It has been found that UV-B can improve photoprotection capacity, however, the process of UV-B mediates photoprotection mechanism is unclear. In this project, tomato seedlings will be selected as testing species, and UV-B will be appropriately applied to induce the formation of photoprotection mechanisms, so as to improve the adaptability of seedlings to dynamic light after transplantation. Firstly, using plant functional structure model and anatomical methods to study UV-B mediating plant phenotype and chloroplast movement, and further explore their effect on light capture under dynamic light, and clarify the response of plant light harvesting mechanism to UV-B. Secondly, based on chlorophyll a fluorescence kinetic method, study UV-B mediating energy dissipation process of photosynthetic apparatus under dynamic light. In the meanwhile, explore the network and key genes of UV-B mediated energy dissipation and UVR8 photoreceptors and downstream signaling partners through transcriptional and proteomics technology. Finally, study the effect of UV-B mediated photoprotection mechanisms on photosynthetic performance of seedlings that is transplanted to greenhouse condition. This study will be helpful to understand the physiological nature of UV-B affecting photosynthesis, and to provide a theoretical basis for horticultural seedling production.