凯氏带是在内皮层细胞早期分化过程中形成的一种疏水性结构，它的存在限制了基本组织和维管组织之间的自由运输，迫使根系有选择性地吸收营养物质和水分，增强了植物对环境的适应性。凯氏带的出现标志着内皮层细胞发育成熟。我们前期的研究结果发现SHR能和来源于维管束的小肽CIF信号协作，对细胞命运进行重编程，促使非内皮层细胞形成完整的有功能的凯氏带。但是SHR与CIFs两种信号通路如何协同调控凯氏带的形成并不清楚，在本申请中，我们将针对这个未知问题进行探讨，研究SHR-CIF信号调控植物内皮层凯氏带形成的机理，并发掘其介导的调控网络。

Casparian strip is an apoplastic diffusion barrier that forms in earlier stage of endodermal differentiation to enhance the adaptability of plants to environment. The presence of Casparian strip restricts the transport of apoplastic water and nutrient into the endodermal cells and vascular tissues, which is used as a marker for a functional endodermis. Our present findings showed that SHR combined with CIF Peptides induces functional Casparian strips in non-endodermal cells. However, it remains unclear how SHR and CIFs regulate functional Casparian strip formation. Here, we will focus on this question revolving the function of SHR and CIFs in endodermal differentiation and explore its molecular mechanism.