The technical development of the primate glottal activities including fossil hominid.

原始人类声门音的起源可以追溯到6500万年前。此时，在第三纪早期就获得了对树栖三维空间的运动适应的树鼩和早期灵长类动物，其功能--形态适应从嗅觉-脑向视觉-脑转变。与此相对应的是，树鼩和早期灵长类动物的喉腔也获得了运动器官等功能性适应。树猴和灵长类动物的喉囊和特化球囊被认为高于树猴、原猴、猴或树栖动物的运动适应结构。也就是说，包括原猴在内的灵长类动物的声门和前肢可能已经完成了向人类声门音形成的进化，而它们各自功能的协同作用主要是在阻气方面进化的。人类声门音形成是在第三纪早期与对树栖生活的运动适应一起获得的功能。这种功能形态学上适应的喉腔本身的持续维持，没有其特化，而是作为一般化的特征，被认为导致了人类的gl. otal音形成功能。

The origin of hominids' glottal tone goes back to 65 million years ago. At that time, treeshrews and early primates, which had' acquired locomotive adaptation of the arboreal three-dimensional space in early Tertiary, made their functions-morphological adaptation from smell-brain to sight-brain. Pararell to this the laryngeal cavity of treeshrews and early primates also acquired functional adaptation such as locomotive 'organ. The laryngeal sac and specialized saccule of treeshews and primates is thought to be higher than the locomotion adaptive structure of treeshrew, prosimian, monkey or age 'of arboreal life. In other words, , the glottis and forelimbs of primates including prosimian have presumably finished anagenesis toward the-formation of humans-' glottal tone, while the synergism of their respective functions was evolving mainly on the air-trapping. Human glottal tone forming for articulate speech is a function aquired together with the locomotive adaptation of the arboreal life in early Tertiary. The persistent maintenance itself of this functionomorphologically adapted laryngeal cavity without its specialization but as generalized character is thought to lead to the human gl, ottal tone forming function.