Studies on the phylogeny of the Pamphiliid sawflies (Hymenoptera)

1 1992 ~ 1994年6、7月主要在北海道采集锯蝇成虫和幼虫，以丰富以往积累的资料。通过比较形态学的研究，发现在脉脉、下颌骨、产卵器形状等方面有13个特征可用于在属水平上分析系统发育。利用MacClade （Maddison & Maddison, 1992）和PAUP （Swofford, 1993）分析了9个恋虫属之间的系统发育关系，并基于简约准则提出了一种关系假设。根据这一假说，Pamphiliinae亚科及其两个组成部落Neurotomini和Pamphiliini是单系的，而Cephalciinae亚科及其一个组成部落Cephalciini则是非单系的研究过程中的部分新发现已经发表在10篇独立的论文中。其中包括对韩国神经瘤属的一种（5种）和Onycholyda（6种，包括1新种）的修订[文献6]，对Pamphilius sulureipes群的一份修订（4个东亚种，包括2个新亚种）[文献8]，对Pamphilius histrio群的一份修订，包括分支分析（11个全北极种，包括1个新种和1个新亚种）[文献10]，东亚大叶虫属（Megaxyela）的修订（3种包括2新种）[文献3]，日本Cephalcia属和Neurotoma属各1新种的描述[文献5,9]，关于新寄主和幼虫筑巢习性的报道[文献1,7]和新分布记录[文献2,4]。

1 Collection of adults and larvae of the Pamphiliid sawflies was made mainly in Hokkaido in June and July, 1992 to 1994, to enrich the previously accumulated material. As a result of comparative morphological studies, 13 characters in the venation, mandible, shape of the ovipositor, etc., have been found useful for analyzing phylogeny at the generic level. Using MacClade (Maddison & Maddison, 1992) and PAUP (Swofford, 1993), phylogenetic relationships among the nine Pamphiliid genera were analyzed and a hypothesis of the relationships was presented based on parsimony criterion. According to this hypothesis, the subfamily Pamphiliinae and its two component tribes, the Neurotomini and Pamphiliini, are monophyletic, whereas the other subfamily Cephalciinae and one of its component tribes, the Cephalciini, are non-monophyletic.2 Part of new findings during the course of the studies heve been published in ten separate papers. They include a revision of Korean species of the genera Neurotoma (five species) and Onycholyda (six species including one new) [reference 6], a revision of Pamphilius sulphureipes group (four East Asian species including two new subspecies) [reference 8], a revision of Pamphilius histrio group, including cladistic analysis (eleven Holarctic species including one new species and one new subspecies) [reference 10], a revision of the Xyelid genus Megaxyela of East Asia (three species including two new) [reference 3], descriptions of one new species each of the genera Cephalcia and Neurotoma from Japan [reference 5,9], reports on new hosts and larval nesting habits [reference 1,7]and new distribution records[reference 2,4].