Malacostracan phylogeny and the evolution of the caridoid facies

软甲纲是一个高度多样化（约40，000种）和高度不同（17目）的类群内并系甲壳动物。虽然解剖复杂性和/或生态的重要性，许多这些高等甲壳类动物是关键的利益，我们的理解发生在软甲纲内的进化转变是阻碍了一个事实，即分类群内的系统发育关系仍然在很大程度上没有得到解决。这并不意味着形态学和分子基础的分析是不一致的。一方面，软甲介类的发生主要方面是有争议的，即使是在形态学的基础上，另一方面，基于分子的分析是相对罕见的。本提案的总体目标是，利用微型CT和数字三维重建等新技术，对各种软甲甲壳类动物的肌肉组织和表皮骨骼进行详细调查，作为新特征和特征状态概念化的基础，并将其添加到现有特征矩阵中。新创建的字符矩阵将是有史以来最广泛的形态矩阵编制软甲藻，并将形成各种系统发育分析的基础上，测试的稳健性软甲藻属。结果将与分子系统发育分析，包括我们自己的基因组分析进行对比。由此获得的软介虫的进化假说将被用来制定和讨论有关进化转变的假说（特别是那些形成本项目的重点）。一个修订和扩展版本的caridoid相假说将提出。

Malacostraca is a highly diverse (numbering around 40,000 species) and highly disparate (17 orders) taxon within paraphyletic crustaceans. Although the anatomical complexity and/or ecological importance of many of these higher crustaceans are of pivotal interest, our understanding of the evolutionary transformations that took place within Malacostraca is impeded by the fact that the phylogenetic relationships within the taxon are still largely unresolved. This does not just mean that morphological and molecular-based analyses are not congruent. For one thing, major aspects of malacostracan phylogeny are controversial even on the basis of morphology alone and for another, molecular based analyses are relatively rare. The general objective of this proposal is a detailed investigation, using new techniques such as micro-CT and digital 3D reconstruction, of the musculature and the cuticular skeleton in various malacostracan crustaceans for use as a basis for the conceptualization of new characters and character states, which will be added to existing character matrices. The newly created character matrix will be the most extensive morphological matrix ever compiled for Malacostraca and will form the basis of various phylogenetic analyses testing the robustness of malacostracan phylogeny. The results will be contrasted with molecular phylogenetic analyses including our own phylogenomic analysis. The malacostracan phylogeny hypothesis thus obtained will be used to formulate and discuss hypotheses about evolutionary transformations (particularly those which form the focus of the present project). A revised and extended version of the caridoid facies hypothesis will be presented.