The technical development of the primate glottal activities including fossil hominid.

原始人声门音的起源可以追溯到6500万年前。与此同时，树鼩和早期灵长类动物在第三纪早期已经获得了对树木三维空间的快速适应，它们的功能形态适应也从嗅觉脑向视觉脑转变。与此同时，树鼩和早期灵长类动物的喉腔也获得了功能上的适应，如运动器官。树树和灵长类动物的喉囊和特殊囊被认为比树树、原猴、猴子或树栖动物的运动适应结构更高。换句话说，包括原猿在内的灵长类动物的声门和前肢可能已经完成了向人类声门音形成的进化，而它们各自功能的协同作用主要是在空气捕获上进化的。人类发声的声门形成是第三纪早期树栖生物的快速适应过程中所获得的一种功能。这种功能形态上适应喉腔的持续维持本身没有其特化，但作为一种普遍的特征，被认为导致了人类的整体音调形成功能。

The origin of hominids' glottal tone goes back to 65 million years ago. At that time, treeshrews and early primates, which had' acquired locomotive adaptation of the arboreal three-dimensional space in early Tertiary, made their functions-morphological adaptation from smell-brain to sight-brain. Pararell to this the laryngeal cavity of treeshrews and early primates also acquired functional adaptation such as locomotive 'organ. The laryngeal sac and specialized saccule of treeshews and primates is thought to be higher than the locomotion adaptive structure of treeshrew, prosimian, monkey or age 'of arboreal life. In other words, , the glottis and forelimbs of primates including prosimian have presumably finished anagenesis toward the-formation of humans-' glottal tone, while the synergism of their respective functions was evolving mainly on the air-trapping. Human glottal tone forming for articulate speech is a function aquired together with the locomotive adaptation of the arboreal life in early Tertiary. The persistent maintenance itself of this functionomorphologically adapted laryngeal cavity without its specialization but as generalized character is thought to lead to the human gl, ottal tone forming function.