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Speciation and diversification of populations based on disjunct distribution of primitive monocotyledons

基于原始单子叶植物间断分布的种群物种形成和多样化

基本信息

批准号：
15570083
负责人：
TAMURA Minoru N.
金额：
$ 2.37万
依托单位：
Osaka City University
依托单位国家：
日本
项目类别：
Grant-in-Aid for Scientific Research (C)
财政年份：
2003
资助国家：
日本
起止时间：
2003 至 2004
项目状态：
已结题

项目摘要

This study was carried out from the respects of phylogeny, phytogeography and taxonomy at specific level and at infraspecific level, focusing especially on the Tofieldiaceae that have been considered one of the most morphologically primitive monocots and are disjunctly distributed, growing in rocky alpine places, alpine meadows, rock faces along rivers, etc., and often on calcareous soils.First, using plastid trnK, trnL and trnL-F sequences and nuclear ITS sequences, we conducted a combined analysis to clarify phylogenetic relationships of the species and infraspecific taxa of Tofieldia and Triantha. Among the three clades that constitute Tofieldia and the clade covering Triantha, each clade turned out to consist of one widely distributed boreal species, with several localized southern species, excepting the clade of Tofieldia calyculata that occupies an isolated phylogenetic position. Further, morphology of leaf apex and leaf margin proved to be of great taxonomic value in Tofieldia.Second, in order to reexamine the taxonomic coherence of Tofieldia nuda, which is disjunctly distributed in eight areas of East Asia, we included individuals from all of the eight distribution areas of T. nuda in the molecular as well as morphological analyses, and proved that T. nuda contains three species and two varieties. The first species corresponds to T. nuda sense stricto, being sister to Tofieldia coccinea. The second species corresponds to Tofieldia furusei, to which we change the status from T. nuda var. furusei. This species is sister to the clade of Tofieldia divergens and Tofieldia thibetica. The third species corresponds to Tofieldia yoshiiana, which was often reduced to T. nuda by former taxonomists. Tofieldia yoshiiana is further divided into three varieties, i.e. var. yoshiiana, var. kanwonensis, comb. nov., and var. hyugaensis, var. nov.

本研究从植物发生学、植物地理学和种下分类学等方面进行了系统的研究，特别是对分布于高山岩石、高山草甸、沿着岩壁等地的、形态上最原始的单子叶植物--托菲尔蕨科进行了研究。首先，利用质体trnK、trnL和trnL-F序列以及核ITS序列，对Tofieldia和Triantha的种和种下类群的系统发育关系进行了分析。在构成Tofieldia的三个分支和分支覆盖Triantha，每个分支原来是由一个广泛分布的北方种，与几个本地化的南方种，除了分支Tofieldia calyculata，占据一个孤立的系统发育位置。其次，为了重新检验裸叶石楠（Tofieldia nuda）在分类学上的一致性，我们选取了分布于东亚8个地区的裸叶石楠（Tofieldia nuda）的8个分布区的所有个体，并对裸叶石楠（Tofieldia nuda）的形态特征进行了研究。nuda的分子生物学和形态学特征，证明T.裸花有三个种和两个变种。第一个种为T. nuda sense stricto，是Tofieldia coccus的姐妹。第二个种对应于Tofieldia furusei，我们将其从T.努达湾古生。该物种是分支Tofieldia divergens和Tofieldia thibetica的姐妹。第三个种对应于Tofieldia yoshiiana，它经常被简化为T。nuda由前分类学家。Tofieldia yoshiiana进一步分为三个变种，即var.吉亚纳湾kanwonensis，comb. nov.，和var。日向变种Nov.