Evaluation of the Ecdysozoa versus Articulata hypothesis on the basis of complete mitochondrial genome data

计划项目在头两年期间的预期目标是评估蜕皮动物与关节动物假说，从而确定节肢动物在原口动物中最可能的位置。随着分子生物学资料的不断增加，节肢动物的起源问题再次引起人们的争论。传统上，大多数形态学数据被视为支持关节动物门假说的环节动物门和节肢动物门之间姐妹群关系的论据。另一方面，分子数据将节肢动物作为姐妹分类单元与部分线虫动物（Cycloneuralia）（支持分类单元Ecdysozoa）并将环节动物与软体动物和触手动物、部分板虫动物和一些小类群统一为Lophotrochozoa。在计划的项目中，将确定总线粒体基因组序列-特别是线虫-并将其与节肢动物，软体动物，环节动物和触手动物的线粒体基因组序列进行遗传学比较，无论是数据库中已有的还是我们或合作研究人员尚未确定的。此外，随着新的数据，特别是那些线虫，一个更详细的图片内的节肢动物的系统发育关系的系统发育数据分析的结果之一。

The envisaged goal of the planned project within the first two years period is the evaluation of the Ecdysozoa versus the Articulata hypotheses and thus to determine the most likely position of the Arthropoda within the Protostomia. Caused by an increasing amount of molecular data, the origin of the Arthropoda is controversely discussed once again. Traditionally most morphological data are viewed as arguments for a sister group relationship between Annelida and Arthropoda favoring the Articulata hypothesis. On the other hand, molecular data place the arthropods rather as the sister taxon to part of the Nemathelmithes (Cycloneuralia) (supporting the taxon Ecdysozoa) and unify annelids together with the molluscs and tentaculates, part of the Plathelminthes and some minor groups to the Lophotrochozoa. Within the planned project total mitochondrial genome sequences -especially of Nemathelminthes- will be determined and phylogenetically compared to mitochondrial genome sequences of Arthropoda, Mollusca, Annelida and Tentaculata either already available in the database or still to be determined by us or cooperating researchers. Furthermore, with the new data, especially those of Nemathelminthes, a much more detailed picture of phylogenetic relationships within the Arthropoda is expected as one of the results of phylogenetic data analyses.