Inferred Phylogeny of Carabid Tribes with Focal Studies on Taxa of Harpalinae: a Synthesis of Morphological and Molecular Data, and a New Classification

该项目将重建步甲(鞘翅目：步甲科)的系统发育，这是小型捕食者的优势类群之一。重点将放在这个大家庭的部落之间的关系上，重点放在最大的亚科Harpalinae内的关系上，自白垩纪以来，这个群体已经辐射到超过25,000个物种。在Harpalinae中，将详细研究基本的翼片石谱系，以及丰富的利生物形态，它们向上生长并辐射到雨林树冠中。将利用大量已发表的记载、新采样的分类群和新颖的特征系统，构建一个来自200多个分类群的成虫和幼虫的形态特征矩阵。28S rDNA和无翼基因的分子序列数据将被收集到这些相同的分类群，通过添加对理解一般宪兵器类进化至关重要的分类群来扩展关于这些基因的现有信息，特别是骆驼蓬碱辐射。该项目不仅将产生关于步甲类进化史的知识，还将首次基于明确和全面的系统发育分析对该科进行分类。

This project will reconstruct the phylogeny of carabid beetles (Coleoptera: Carabidae), one of the dominant groups of small predators. The focus will be on relationships of the tribes of this large family, with a concentration on the relationships within the largest subfamily, the Harpalinae, a group which has radiated into over 25,000 species since the Cretaceous. Within the Harpalinae, the basal pterostichite lineages will be examined in detail, as will the abundant lebiomorphs, which have ascended vegetation and radiated into the rainforest canopy. A matrix of morphological characters from both adults and larvae of over 200 taxa will be constructed that draws on plentiful published accounts, newly sampled taxa and novel character systems. Molecular sequence data for 28S rDNA and the wingless gene will be gathered for these same taxa, expanding existing information about these genes by adding taxa critical to understanding carabid evolution in general and harpaline radiation in particular. The project will yield not only knowledge of the evolutionary history of carabids, but a classification of the family based for the first time on an explicit, and comprehensive, phylogenetic analysis.