Systematics and Evolution of the tribe Arabideae with special emphasis on the genus Arabis (Brassicaceae)

广泛定义的属南芥已修订系统只有在少数几个部分。一些以前被分配到南芥属的物种现在被限制在Turnalus，Pseudoturrits，Boechera或Fourraea等属。其他属，如拟南芥/豆蔻属或Borodinia过去被错误地归入南芥属。与此相反，Rhammotophyllum、Pachyneurum或Botschantzevia等属可能最好归入修订后的Arabideae族。上面提到的一些属不是阿拉伯芥科的成员，考虑到实际的知识，实际上有70到140种阿拉伯芥属的物种，以及大约370种Draba属及其最近的亲属可能代表了阿拉伯芥科的核心。我们的初步研究表明，葶苈属及其分离种大多是单系的，而南芥属的种则是一个帕拉和多系的种集合。 物种丰富和广泛分布的物种复合体，如Arabis hirsuta-aggregate在过去也被错误地解释，很明显，杂交，多倍化和网状化在许多物种的进化史中起着重要作用。 在这个项目中，我们的目标是分析目前广泛定义的属阿拉伯芥在整个部落的背景下的系统学和进化。我们将重建整个部落的进化史，特别强调更重要和分布更广的物种复合体，如阿拉伯芥的hirsuta聚集，红花北美阿拉伯芥和阿拉伯芥高山组发挥越来越大的作用，在发展和进化研究的模型的进化历史和地理。南芥属植物广泛分布于北半球的高山和山地生境，是研究适应性性状进化的理想系统。

The broadly defined genus Arabis has been revised systematically only in few parts. Some species previously assigned to Arabis are now confined to genera such as Turritis, Pseudoturrits, Boechera or Fourraea. Other genera such as Arabidopsis/Cardaminopsis or Borodinia have been erroneously placed into Arabis in the past. In contrast genera such as Rhammotophyllum, Pachyneurum or Botschantzevia might be best placed in a revised tribe Arabideae. Some of the above mentioned genera are not members of tribe Arabideae, and considering the actual knowledge actually 70 to 140 Arabis species together with approximately 370 species of the genus Draba and its closest relatives might represent the core of tribe Arabideae. Our preliminary work has demonstrated that Draba and its segregates are mostly monophyletic, but in contrast, Arabis species appeared as a largely para- und polyphyletic set of species. Species-rich and widely distributed species complexes such as the Arabis hirsuta-aggregate have been also wrongly interpreted in the past, and it is apparent that hybridization, polyploidization and reticulation play an important role in the evolutionary history of many species. In this project we aim to analyze the systematics and evolution of the currently broadly defined genus Arabis in the context of the whole tribe Arabideae. We will reconstruct the phylogeny of the whole tribe with special emphasis on the evolutionary history and phylogeography of the more important and wider distributed species complexes such as the Arabis hirsuta-aggregate, red-flowering North American Arabis and the Arabis alpina group playing an increasing role as model in developmental and evolutionary research. Representatives of the genus Arabis are distributed in all north-hemispherical alpine to montane habitats and provide an ideal system to study the evolution of adaptive characters and traits.