Evolution of a giant genus (II): A phylogenetic case study within Peperomia (Piperaceae)

在过去的两年中，已取得重大进展，对所产生的数据和主要的系统和分类问题的解决方案，在物种丰富的属胡椒。此外，序列数据已经产生，形态特征已被评分，以跟踪字符的演变。前两年的各项预定任务已全部完成。虽然我们已经发表了几篇论文，记录了所取得的进展，但还需要两年时间来完成现有的数据集，分析和解释数据，并发表主要论文。具体而言，我们将重点关注：i）上一次实地考察的序列材料，重点是更古老的主要附生属的（陆生）物种，ii）用来自这些收集物的加入物和从合作伙伴获得的材料扩大骨干系统发育，iii）评估所产生的数据的系统发育含义，特别是在分子进化方面（如速率异质性）以及系统和分类学意义，iv）增加生物地理学方法，通过对地生物种的几乎完全的采样以及对整个属的代表性的世界范围的采样，最后但并非最不重要的五）我们将进行祖先地区重建和分子年代测定分析，提供尽可能完整的这一群体的演变情况。

In the last two years, major progress has been made towards a synthesis of generated data and solution of major systematic and taxonomic issues in the species-rich genus Peperomia. Furthermore, sequence data have been generated and morphological traits have been scored to trace character evolution. All scheduled tasks of the first two years have been accomplished. Although we already published several papers documenting the progress made, two additional years are needed to complete the existing datasets, to analyze and interpret the data as well as to publish the main papers. In detail we will focus on: i) sequence material from the last field trip which focused on the more ancestral (terrestrial) species of the mainly epiphytic genus, ii) enlarging the backbone phylogeny with accessions from these collections and material obtained from cooperation partners, iii) evaluate the phylogenetic implication of the generated data, especially with respect to molecular evolution (e.g. rate heterogeneity) as well as systematic and taxonomic implications, iv) adding a biogeographical approach, enabled by an almost complete sampling of the geophytic species as well as a representative worldwide sampling for the whole genus, and last but not least v) we will perform ancestral area reconstructions and molecular dating analyses, providing a picture as complete as possible of the evolution of this group.