Biotic history has accelerated under anoxic environments - Paradox of Foraminifera evolution can solve through field observation, laboratory culture and genomic analyses.

我们假设活跃的微生物共生可能会增强海洋原生生物的进化。当海洋缺氧环境占主导地位时，与微生物的共生关系适合在危机期间生存。对于真核单细胞生物来说，这场危机并不是真正的危机，因为它们可能通过各种类型的共生来积极适应这种缺氧。我们通过实地调查、准确的环境评估、基因组学、精确的实验室培养和生物地球化学测量来检验这一假设。在相模湾、下田湾、太平洋（八户附近）、上越岛的波子池等地进行了实地研究。研究结果通过 29 篇论文、42 篇演讲等形式向科学界报告。同时，这些知识通过互联网、研讨会、新闻发布等场合立即转移到社会。

We hypothesized that the evolution of marine protists might been enhanced by active microbial symbiosis. The symbiotic relationships with microbes were suitable to survive during the crisis when the anoxic environments were dominant in the ocean. The crisis is not really crisis for eukaryotic unicellular organisms because they may potentially positively adaptable to this anoxia by various types of symbiosis. We have tested the hypothesis through field surveys, accurate environment assessments, genomics, precise laboratory culture and biogeochemical measurement. Field studies were carried out at Sagami Bay, Shimoda Bay, Pacific Ocean (off Hachinohe), Namako-ike lake of Kamikoshiki-jima island, etc. The results are reported through 29 papers, 42 presentations and others for science community. Meanwhile, the knowledge has immediately transferred to society by Internet, seminars, press release and other occasions.