Ecological studies on relationship between related species and species structure of macro-soil-invertebrates, especially insects

采伐、种植或间伐后地甲群落结构的变化：开阔地或林缘物种多样性增加，群落结构的年际变化和季节变化也较明显。近缘种间生活史的分化：在一些亚科和属中，可以推测繁殖季节向春季繁殖者或秋季繁殖者的分化是平行发生的。杂交区的实地调查和Ohomopterus亚属(Carabus，Carabidae)稀有物种的杂交试验表明：由于杂交个体的直接代价，两种间的杂交维持了毗邻分布格局并允许部分基因在物种间流动。这一结果暗示了该亚属的网状进化。

The change of community structure of ground beetles following clearcutting and planting or thinning at forests : the species diversity increased at the open area or the edge of forest, and the annual change and the seasonal change of community structure were also recognized considerably.The differentiation of life history between related species belonging to Carabidae : in some subfamilies and genera, it was inferred that the differentiation of reproductive season to spring breeder or to autumn breeder occurred parallelly.The field surveys of hybrid zone and crossing experiments between parapatric species belonging to subgenus Ohomopterus (genus Carabus, Carabidae) : the interspecific hybridization between two species maintain the parapatric distribution pattern because of the direct cost to hybridizing individuals, and allow partial gene flow across the species. This result suggests the reticulate evolution in this subgenus.