Ecological forces driving the loss of photosynthesis in phytoplankton: Molecular and morphological adaptations to heterotrophy

营养策略的分化（光养、兼养、食养）是生物体间最明显的功能差异之一，并强烈影响着碳和元素的流动。所有主要的藻类谱系都包括许多“无色藻类”的例子。我们解决的分子和形态的适应异养金藻作为模式生物的光合能力丧失的生态原因得出结论。我们将通过一系列的实验室实验进一步测试异养和混合营养型金藻在不同条件下的竞争强度。该项目汇集了各工作组的专门知识，涵盖藻类多样性和藻类发生、超微结构和细胞生物学以及基因组和转录组分析，特别是藻类色素的演变。这是进一步独立，但密切相关的建议，同时提交的美国。盖德基它将提供基本的见解背后的（多重）分离到光养和异养谱系的情况，并阐明有利于这种分化的环境条件。

The differentiation of nutritional strategies (phototrophic, mixotrophic, phagotrophic) is one of the most obvious functional differences among organisms and strongly influences the carbon and element flows. All major algal lineages comprise numerous examples of "colourless algae". We address the molecular and morphological adaptations to heterotrophy using chrysophytes as model organisms to conclude on ecological causes of the loss of phototrophic abilities. We will further test the competitive strength of heterotrophic and mixotrophix chrysophytes under different conditions in a series of laboratory experiments. The project brings together the expertise of working groups covering algal diversity and phylogeny, ultrastructure and cell biology, and genome and transcriptome analyses with special reference to the evolution of photopigments. It is further independent but closely linked to a proposal submitted in parallel by U. Gaedke. It will provide basic insights into the scenarios behind the (multifold) separation into phototrophic and heterotrophic lineages and elucidate environmental conditions favouring this differentiation.