Morphological differentiation and life history of sympatric and allopatric dryopteroid ferns

1992-1993年期间进行了以下两项主要工作。(1)对美国西北部和日本最北部20种蕨类植物的叶片异速生长和相对蛋白质组成进行了调查。(2)对北海道蕨类植物的微分布格局和频率进行了连续调查。在日本最北部和美国西北部的蕨类植物中，相对蛋白质组成与叶片异速形态没有相关性。可能产生自然杂交的同域蕨类植物的相对蛋白质组成有相同的趋势。另一方面，同感蕨类植物在叶片异速生长方面与东京毛蕨（Dryopteris tokyoensis）、monticola、cristata和goldiana存在明显差异。尺度分析包括两种方法，一种是从同一区域的不同分辨率(a)推导出的，另一种是从局部到全球尺度的不同范围大小的相同数量的样方(B)推导出的。这里介绍第二种技术(B)。每个物种以频率（%）表示其各自的轨迹，即脆弱囊蝇在北半球尺度上频率较高，而在北海道尺度上频率较低。这些波动轨迹与尺度分析可能受到不同类型因素的影响，如历史陆地迁移和最近的气候条件。

Two main works were conducted during 1992-1993 as follows. (1) Leaf allometry and relative protein composition were surveyed among 20 fern species from north-western America and northernmost Japan. (2) And micro-distribution patterns and frequency were continuously surveyed on Hokkaido pteridophytes.No correlation was found between relative protein composition and leaf allometry patterns on dryopteroid ferns from northernmost Japan and north-western America. There was a tendency that sympatric ferns with possibile production of natural hybrid showed same pattern of the relative protein composition. On the other hand, the sympatic ferns showed clear difference of leaf allometry between Dryopteris tokyoensis and D.monticola, and D.cristata and D.goldiana.Spatial distribution patterns of some cool temperate ferns found in northernmost Japan were quantified by the scaling analysis. Scaling analysis includes two ways, one is derived from different resolution in a same area (A) and another is derived from same number of quadrats in different size of scope (B) from local to global scales. Here introduced the second technique (B). Each species shows its own trajectory as frequencies(%), i.e., Cystopteris fragilis shows the high frequency in northern hemisphere scale however does the low frequency in Hokkaido scale. These fluctuating trajectories with scaling analysis might be influenced by the different kind of factors, such as historical land migration and recent climatic conditions.