Ecological studies on relationship between related species and species structure of macro-soil-invertebrates, especially insects

森林采伐和间伐后地甲虫群落结构的变化：开阔区和林缘的物种多样性增加，群落结构的年际变化和季节变化也得到了较好的认识。卡拉贝科近缘种生活史的分化：在某些亚科和属中，推测繁殖季节向春季繁殖体或秋季繁殖体的分化是平行发生的。本研究通过对大翅猴亚属（Carabus属，carabides科）异翅猴亚属异翅猴种杂交带的野外调查和杂交实验发现：两种间的种间杂交由于对杂交个体的直接代价而维持了异翅猴的分布格局，并允许部分基因在种间流动。这一结果表明该亚属的网状进化。

The change of community structure of ground beetles following clearcutting and planting or thinning at forests : the species diversity increased at the open area or the edge of forest, and the annual change and the seasonal change of community structure were also recognized considerably.The differentiation of life history between related species belonging to Carabidae : in some subfamilies and genera, it was inferred that the differentiation of reproductive season to spring breeder or to autumn breeder occurred parallelly.The field surveys of hybrid zone and crossing experiments between parapatric species belonging to subgenus Ohomopterus (genus Carabus, Carabidae) : the interspecific hybridization between two species maintain the parapatric distribution pattern because of the direct cost to hybridizing individuals, and allow partial gene flow across the species. This result suggests the reticulate evolution in this subgenus.