What factors govern the occurrence of highly developed territorial behavior termed barricade building in Ilyoplax pusilla (Crustacea ; Brachyura)?

本文从以下几个方面探讨了影响滩涂蟹类特有的地域性行为--障碍物发生的因素：1.外界因素对障碍物出现频率的影响在繁殖季节，障碍物的出现频率增加，但与潮汐关系不明显。2.障碍建筑与交配成功率的关系结对雄性与非结对雄性之间的比较表明，结对雄性比非结对雄性建造障碍的频率更高。3.土结构回避倾向的种间比较利用…比较粉刺石首鱼和其他两种斑腿类对人工土结构的反应4.模型构建在一个模型中，分别给出了路障(建、不建、建小)和路障(毁、不毁、毁小)三种策略，并研究了它们频率的进化动态。该模型表明，路障的成本应该与频率相关，以解释为什么只有部分人口成员修建路障。然而，均衡只是中性稳定的，随机遗传漂移应该会在没有障碍的情况下改变系统。引入频率依赖策略，根据路障的频率来摧毁路障，使路障与中间路障的频率保持稳定。在另一个模型中，用动态规划方法研究了生活史策略。在ESS中，只在繁殖期修建路障，这与现场数据重合较少

Factors governing the occurrence of unique territorial behavior termed barricade building by the brachyuran crab Ilyoplax pusilla (family Dotilllidae) living on tidal flat, have been explored for the following aspects.1.Influence of external factors on frequency of barricade buildingThe frequency of barricade building increased in the breeding season, but was not correlated with the tidal regime. The frequency was also not related with the population density or with difference of the substratum condition.2.Relationship of barricade building with mating successComparison of occurrence of barricade building between pairing males and non-pairing males revealed that the former made barricade building more frequently than the latter. This finding suggests that barricade functions advantageously for pair formation.3.Interspecific comparison of earthen-structure avoiding tendencyComparison of response to artificial earthen structure between Ilyoplax pusilla and other two dotillid species with … More out barricade building behavior, revealed that I.pusilla less frequently broke the structure or avoided it more frequently than the other two species.4.Model constructionIn one model, three strategies each for barricader (build, do not build, build to smaller) and barricadee (destroy, do not destroy, destroy to smaller) are assigned, and the evolutionary dynamics of their frequency has been investigated. The model shows that cost by barricade should be frequency dependent to explain why only part of population members build barricades. However, the equilibrium is only neutrally stable and random genetic drift should turn the system with no barricade. Introduction of frequency depended strategy, destroy barricade depending on their frequency, made the equilibrium with intermediate barricade frequency stable. In the other model, life history strategies have been investigated by dynamic programming method. In ESS, barricade is built only in the breeding season, which coincides with field data Less