Diversity, adaptation and coevolution in snail-feeding carabid beetles and land snails

为了了解陆地蜗牛区系与Carabina亚族中与蜗牛喂养相关的形态学差异（大头症与窄头症）之间的关系，我们在中国，韩国，亚美尼亚和摩洛哥进行了实地研究。在中国内陆地区，尤其是不同的食螺Carabina物种和陆生蜗牛发生。大头石首鱼物种出现在栖息地窝藏小尺寸的蜗牛丰富。在栖息地，宏观和狭头的Carabina物种发生，壳形态的缓贝蜗牛显着多样化，这表明一个适应性辐射由于强大的捕食压力，从蜗牛喂养的捕食者。

To understand how the land snail fauna is related with the divergence of morphology related to snail feeding (macrocephalism vs. stenocephalism) in the subtribe Carabina, we conducted field studies in China, Korea, Armenia, and Morocco. In the inland region of China, especially, diverse snail-feeding Carabina species and land snails occur. Macrocephalic Carabina species occur in habitats harboring small-sized bradybaenid snails abundantly. In habitats where both macro- and stenocephalic Carabina species occur, the shell morphology of bradybaenid snails was markedly diversified, suggesting an adaptive radiation due to strong predation pressure from snail-feeding predators.